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Triassurus
''Triassurus'' is an extinct genus of amphibian, and the oldest member of Caudata (salamanders and close relalatives). It is known from the Middle to Upper Triassic (Ladinian-Carnian) aged Madygen Formation in Kyrgyzstan. The type species is ''T. sixtelae''. Description The holotype specimen is a partial and poorly preserved skeleton (PIN-2584/10), including the skull. Holotype specimen had tiny skull just long, but it was probably a larva: the neural arches of the vertebrae were still paired and no vertebral centers show any degree of ossification. The skull closely resembles that of current salamanders, especially as regards the long space in the joint of the maxillary bones and the lack of bone connection between the pterygoid and the maxilla. There were about 20 presacral vertebrae, while the legs were small. In 2020, a more complete adult specimen (FG 596/V/20) with long skull was described. Classification ''Triassurus'' was first described in 1978, based on a fossil found ...
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Madygen Formation
The Madygen Formation (Russian language, Russian: Madygen Svita) is a Middle Triassic, Middle–Late Triassic (Ladinian–Carnian) geologic formation and lagerstätte in the Batken Region, Batken and Osh Regions of western Kyrgyzstan, with minor outcrops in neighboring Tajikistan and Uzbekistan. The conglomerate (geology), conglomerates, sandstones and mudstones of the thick formation were deposited in terrestrial lacustrine, alluvial, fluvial and river delta, deltaic depositional environment, environments. The formation, extending across the Fergana Valley and Fergana Range, is unique for Central Asia, as it represents one of the few known continental deposits and the Madygen Formation is renowned for the preservation of more than 20,000 fossil insects, making it one of the richest Triassic lagerstätten in the world. Other vertebrate fossils as fish, amphibians, reptiles and synapsids have been recovered from the formation too, as well as minor fossil flora. The lake sediments ...
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Caudata
The Caudata are a group of amphibians containing the extant salamanders (the order Urodela) and all extinct species of amphibians more closely related to salamanders than to frogs. They are typically characterized by a superficially lizard-like appearance, with slender bodies, blunt snouts, short limbs projecting at right angles to the body, and the presence of a tail in both larvae and adults. Disagreement exists between different authorities as to the definition of the terms "Caudata" and "Urodela". Some maintain that Urodela should be restricted to the crown group, with Caudata being used for the total group. Others restrict the name Caudata to the crown group and use Urodela for the total group. The former approach seems to be most widely adopted and is used in this article. Evolution The origins and evolutionary relationships between the three main groups of amphibians ( apodans, urodeles and anurans) is a matter of debate. A 2005 molecular phylogeny, based on rDNA a ...
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Late Triassic
The Late Triassic is the third and final epoch (geology), epoch of the Triassic geologic time scale, Period in the geologic time scale, spanning the time between annum, Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch and followed by the Early Jurassic Epoch. The corresponding series (stratigraphy), series of rock beds is known as the Upper Triassic. The Late Triassic is divided into the Carnian, Norian and Rhaetian Geologic time scale, ages. Many of the first dinosaurs evolved during the Late Triassic, including ''Plateosaurus'', ''Coelophysis'', ''Herrerasaurus'', and ''Eoraptor''. The Triassic–Jurassic extinction event began during this epoch and is one of the five major mass extinction events of the Earth. Etymology The Triassic was named in 1834 by Friedrich August von Namoh, Friedrich von Alberti, after a succession of three distinct rock layers (Greek meaning 'triad') that are widespread in southern Germany: the lower Buntsandstein (colourful ...
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Synapomorphy And Apomorphy
In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to have evolved in their most recent common ancestor. ) In cladistics, synapomorphy implies homology. Examples of apomorphy are the presence of erect gait, fur, the evolution of three middle ear bones, and mammary glands in mammals but not in other vertebrate animals such as amphibians or reptiles, which have retained their ancestral traits of a sprawling gait and lack of fur. Thus, these derived traits are also synapomorphies of mammals in general as they are not shared by other vertebrate animals. Etymology The word —coined by German entomologist Willi Hennig—is derived from the Ancient Greek words (''sún''), meaning "with, together"; (''apó''), meaning "away from"; and (''morphḗ''), meaning "shape, form". Determining apom ...
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Fossils Of Kyrgyzstan
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in amber, hair, petrified wood and DNA remnants. The totality of fossils is known as the ''fossil record''. Though the fossil record is incomplete, numerous studies have demonstrated that there is enough information available to give a good understanding of the pattern of diversification of life on Earth. In addition, the record can predict and fill gaps such as the discovery of ''Tiktaalik'' in the arctic of Canada. Paleontology includes the study of fossils: their age, method of formation, and evolutionary significance. Specimens are sometimes considered to be fossils if they are over 10,000 years old. The oldest fossils are around 3.48 billion years to 4.1 billion years old. Early edition, published online before print. The ob ...
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Crown Group
In phylogenetics, the crown group or crown assemblage is a collection of species composed of the living representatives of the collection, the most recent common ancestor of the collection, and all descendants of the most recent common ancestor. It is thus a way of defining a clade, a group consisting of a species and all its extant or extinct descendants. For example, Neornithes (birds) can be defined as a crown group, which includes the most recent common ancestor of all modern birds, and all of its extant or extinct descendants. The concept was developed by Willi Hennig, the formulator of phylogenetic systematics, as a way of classifying living organisms relative to their extinct relatives in his "Die Stammesgeschichte der Insekten", and the "crown" and "stem" group terminology was coined by R. P. S. Jefferies in 1979. Though formulated in the 1970s, the term was not commonly used until its reintroduction in 2000 by Graham Budd and Sören Jensen. Contents of the crow ...
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Stem-group
In phylogenetics, the crown group or crown assemblage is a collection of species composed of the living representatives of the collection, the most recent common ancestor of the collection, and all descendants of the most recent common ancestor. It is thus a way of defining a clade, a group consisting of a species and all its extant or extinct descendants. For example, Neornithes (birds) can be defined as a crown group, which includes the most recent common ancestor of all modern birds, and all of its extant or extinct descendants. The concept was developed by Willi Hennig, the formulator of phylogenetic systematics, as a way of classifying living organisms relative to their extinct relatives in his "Die Stammesgeschichte der Insekten", and the "crown" and "stem" group terminology was coined by R. P. S. Jefferies in 1979. Though formulated in the 1970s, the term was not commonly used until its reintroduction in 2000 by Graham Budd and Sören Jensen. Contents of the crown ...
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Dissorophoidea
Dissorophoidea is a clade of medium-sized, temnospondyl amphibians that appeared during the Moscovian in Euramerica, and continued through to the Late Permian and the Early Triassic of Gondwana. They are distinguished by various details of the skull, and many species seem to have been well adapted for life on land. Dissorophoid diversity was highest in the Permian; some of the more diverse families within the group include Dissorophidae (toad-like amphibians with armored scutes along their backbone), Trematopidae (terrestrial predators with large triangular skulls), and Branchiosauridae (small neotenic amphibians with large external gills). The small Permo-Carboniferous Micromelerpetontidae are another example of neotenic dissorophoids. Many small dissorophoids with short rounded skulls were historically known as "amphibamids"; in 2018, the name Amphibamiformes was erected for a clade equivalent to the broad historical definition of "Amphibamidae". Since 2008, a consensus ...
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Scapula
The scapula (: scapulae or scapulas), also known as the shoulder blade, is the bone that connects the humerus (upper arm bone) with the clavicle (collar bone). Like their connected bones, the scapulae are paired, with each scapula on either side of the body being roughly a mirror image of the other. The name derives from the Classical Latin word for trowel or small shovel, which it was thought to resemble. In compound terms, the prefix omo- is used for the shoulder blade in medical terminology. This prefix is derived from ὦμος (ōmos), the Ancient Greek word for shoulder, and is cognate with the Latin , which in Latin signifies either the shoulder or the upper arm bone. The scapula forms the back of the shoulder girdle. In humans, it is a flat bone, roughly triangular in shape, placed on a posterolateral aspect of the thoracic cage. Structure The scapula is a thick, flat bone lying on the thoracic wall that provides an attachment for three groups of muscles: intrinsic, e ...
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Squamosal Bone
The squamosal is a skull bone found in most reptiles, amphibians, and birds. In fishes, it is also called the pterotic bone. In most tetrapods, the squamosal and quadratojugal bones form the cheek series of the skull. The bone forms an ancestral component of the dermal roof and is typically thin compared to other skull bones. The squamosal bone lies ventral to the temporal series and otic notch, and is bordered anteriorly by the postorbital. Posteriorly, the squamosal articulates with the quadrate and pterygoid bones. The squamosal is bordered anteroventrally by the jugal and ventrally by the quadratojugal. Function in reptiles In reptiles, the quadrate and articular bones of the skull articulate to form the jaw joint. The squamosal bone lies anterior to the quadrate bone. Anatomy in synapsids Non-mammalian synapsids In non-mammalian synapsids, the jaw is composed of four bony elements and referred to as a quadro-articular jaw because the joint is between the art ...
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Parietal Bone
The parietal bones ( ) are two bones in the skull which, when joined at a fibrous joint known as a cranial suture, form the sides and roof of the neurocranium. In humans, each bone is roughly quadrilateral in form, and has two surfaces, four borders, and four angles. It is named from the Latin ''paries'' (''-ietis''), wall. Surfaces External The external surface [Fig. 1] is convex, smooth, and marked near the center by an eminence, the parietal eminence (''tuber parietale''), which indicates the point where ossification commenced. Crossing the middle of the bone in an arched direction are two curved lines, the superior and inferior temporal lines; the former gives attachment to the temporal fascia, and the latter indicates the upper limit of the muscular origin of the temporal muscle. Above these lines the bone is covered by a tough layer of fibrous tissue – the epicranial aponeurosis; below them it forms part of the temporal fossa, and affords attachment to the temporal mu ...
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