Torosteus
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Torosteus
''Torosteus'' is an extinct genus of arthrodire placoderm from the Early Frasnian stage of the Late Devonian period. Fossils are found in the Kimberley region of Australia. Description ''Torosteuss body size and morphology are similar to ''Incisoscutum'' and ''Compagopiscis'', suggesting a possible pelagic lifestyle, although they were on different trophic levels of their ecosystem. Bite force analysis has suggested that it was an active predator, and fossils have been found in numbers, suggesting possible schooling behavior. Phylogeny ''Torosteus'' was originally classified as a member of the family Plourdosteidae. However, phylogenetic analysis later found Plourdosteidae to be an invalid grouping, and the family was dismissed. ''Torosteus'' is now considered to be a member of the superfamily Incisoscutoidea, which belongs to the clade Coccosteomorphi, one of the two major clades within Eubrachythoraci. The cladogram below shows the phylogeny A phylogenetic tree or phyloge ...
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Incisoscutoidea
Incisoscutoidea is an extinct superfamily of arthrodire placoderms that lived during the Devonian period. Phylogeny Eubrachythoraci is divided into the sister clades Pachyosteomorphi and Coccosteomorphi, the latter of which can be further sub-divided into the two sister superfamilies Coccosteoidea and Incisoscutoidea, as shown in the cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ... below: References {{Taxonbar, from=Q33189516 Arthrodires ...
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Coccosteomorphi
Coccosteomorphi is an extinct clade of arthrodire placoderms within the Eubrachythoraci (of the suborder Brachythoraci), armored fish most diverse during the Devonian. Most are considered to be pelagic (open ocean) long-distance swimmers, leading to their widespread distribution beginning from at least the Middle Devonian period. Phylogeny Coccosteomorphi is the sister taxon to Pachyosteomorphi, which together are the two main sub-clades of Eubrachythoraci. Coccosteomorphi can be further sub-divided into Coccosteoidea and Incisoscutoidea, as shown in the cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ... below: References Arthrodires {{Placoderm-stub ...
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Incisoscutum
''Incisoscutum'' is an extinct genus of arthrodire placoderm from the Early Frasnian Gogo Reef, from Late Devonian Australia. The genus contains two species ''I. ritchiei'', named after Alex Ritchie, a ichthyology, palaeoichthyologist and senior fellow of the Australian Museum, and ''I. sarahae'', named after Sarah Long, daughter of its discoverer and describer, John A. Long. The genus is important in the study of early vertebrates as well-preserved fossilized embryos have been found in female specimens and ossified pelvic claspers found in males. This shows that viviparity and internal fertilization was common amongst these primitive jawed vertebrates, which are outside the crown group Gnathostomata. In a study of fossil remains, comparison of the ontogeny of fourteen dermal plates from ''Compagopiscis, Compagopiscis croucheri'' and the more derived species ''Incisoscutum ritchiei'' suggested that lengthwise growth occurs earlier in the ontogeny than growth in width, and that di ...
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Compagopiscis
''Compagopiscis'' is an extinct genus of placoderm known from the Gogo Formation. It lived in the Upper Devonian (Frasnian stage) of Western Australia. The genus is monotypic, with its only species being ''Compagopiscis croucheri''. Description ''Compagopiscis'' is one of the earliest known vertebrates to have teeth. An analysis of its bite force suggests that it was a generalist predator, having a diverse diet. Fossils have been found in numbers, suggesting possible schooling behavior. Phylogeny ''Compagopiscis'' was originally classified as a member of the family Plourdosteidae. However, phylogenetic analysis later found Plourdosteidae to be an invalid grouping, and the family was dismissed. ''Torosteus'' is now considered to be a member of the superfamily Incisoscutoidea, which belongs to the clade Coccosteomorphi, one of the two major clades within Eubrachythoraci. The cladogram below shows the phylogeny A phylogenetic tree or phylogeny is a graphical representati ...
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Dickosteus Threiplandi
''Dickosteus'' is an extinct genus of coccosteid arthrodire placoderm from the Late Eifelian to Early Givetian stages of the Middle Devonian period. Fossils are found in Orkney and Caithness, Scotland. It was a small placoderm with a total body length of . It is one of the few placoderms for which complete bodies are known. Phylogeny ''Dickosteus'' is a member of the family Coccosteidae, which belongs to the clade Coccosteomorphi, one of the two major clades within Eubrachythoraci. The cladogram below shows the phylogeny A phylogenetic tree or phylogeny is a graphical representation which shows the evolutionary history between a set of species or Taxon, taxa during a specific time.Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, M ... of ''Dickosteus'': References {{Taxonbar, from=Q112327258, from2=Q55762836 Coccosteidae Givetian life Fossils of Scotland Fish described in 1963 ...
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Coccosteus Cuspidatus
''Coccosteus'' (from , 'berry' and 'bone') is an extinct genus of arthrodire placoderm from the Devonian period. Its fossils have been found throughout Europe and North America. The majority of these have been found in freshwater sediments, though such a large range suggests that they may have been able to enter saltwater. It was a small placoderm, with ''Coccosteus cuspidatus'' measuring long. Description Like all other arthrodires, ''Coccosteus'' had a joint between the armor of the body and skull. It also had an internal joint between its neck vertebrae and the back of the skull, allowing for the mouth to be opened even wider. Along with the longer jaws, this allowed ''Coccosteus'' to feed on fairly large prey. The up-and-down movement of the skull also allowed for more water to be pumped through the gills. Possibly, the creature supplemented its diet with organic material filtered from mud using the gills. As with all other arthrodires, ''Coccosteus'' had bony dental pla ...
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Millerosteus Minor
''Millerosteus'' is an extinct genus of coccosteid arthrodire placoderm from the Early Givetian stage of the Middle Devonian period. Fossils are found in Orkney and Caithness, Scotland. It was a small placoderm with a body length of . ''Millerosteus'' is one of the few arthrodires known from specimens preserving the entire skeleton. Phylogeny ''Millerosteus'' is a member of the family Coccosteidae, which belongs to the clade Coccosteomorphi, one of the two major clades within Eubrachythoraci. The cladogram below shows the phylogeny A phylogenetic tree or phylogeny is a graphical representation which shows the evolutionary history between a set of species or Taxon, taxa during a specific time.Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, M ... of ''Millerosteus'': References {{Taxonbar, from=Q112514447, from2=Q97371187 Coccosteidae Givetian life Fossils of Scotland Fish described in 1959 Arthrodire genera ...
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Coccosteidae
Coccosteidae is a family (biology), family of arthrodire placoderms from the Early to Late Devonian. Fossils appear in various strata in Europe, North America and China. Phylogeny Coccosteidae belongs to the larger clade Coccosteomorphi, which together with its sister clade Pachyosteomorphi forms the group Eubrachythoraci. The phylogeny of Coccosteidae can be shown in the cladogram below: Genera ''Belgiosteus'' A genus of very large coccosteids. Species are found in Middle Devonian Belgium and China. ''Clarkosteus'' ''Coccosteus'' The type genus of the family. Numerous species are found in Middle to Upper Devonian strata throughout Europe and parts of North America. ''Dickosteus'' ''Jiuchengia'' The earliest known coccosteid from Late Emsian Yunnan province, China. It is distinguished from other coccosteids by having an elongated occipital. ''Livosteus'' A genus of very large coccosteids known from Middle to Late Devonian strata of Eastern Europe. ''Millerosteus'' A genus ...
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Coccosteoidea
Coccosteoidea is an extinct superfamily of arthrodire placoderms that lived during the Devonian period. Phylogeny Eubrachythoraci is divided into the sister clades Pachyosteomorphi and Coccosteomorphi, the latter of which can be further sub-divided into the two sister superfamilies Coccosteoidea and Incisoscutoidea, as shown in the cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ... below: References {{Taxonbar, from=Q30971248 Arthrodires ...
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Eubrachythoraci
Eubrachythoraci is an extinct clade of arthrodire placoderms within the suborder Brachythoraci, armored fish most diverse during the Devonian. Most are considered to be pelagic (open ocean) long-distance swimmers, leading to their widespread distribution beginning from at least the Middle Devonian period. Phylogeny Brachythoraci is divided into the large derived clade Eubrachythoraci and several basal groups: Buchanosteoidea, Homosteidae, and Holonematidae. (Although Holonematidae's membership in Brachythoraci is disputed.) Eubrachythoraci is then further divided into the sub-clades Coccosteomorphi and Pachyosteomorphi, the latter of which can be further sub-divided into Aspinothoracidi and Dunkleosteoidea, as shown in the cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it doe ...
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Placoderm
Placoderms (from Ancient Greek πλάξ [''plax'', ''plakos''] 'Plate (animal anatomy), plate' and δέρμα [''derma''] 'skin') are vertebrate animals of the class (biology), class Placodermi, an extinct group of prehistoric fish known from Paleozoic fossils during the Silurian and the Devonian geological period, periods. While their endoskeletons are mainly cartilaginous, their head and thorax were covered by articulated armour (zoology), armoured plates (hence the name), and the rest of the body was scale (zoology), scaled or naked depending on the species. Placoderms were among the first jawed fish (their fish jaw, jaws likely Evolution, evolved from the first pair of gill arches), as well as the first vertebrates to have true tooth, teeth. They were also the first fish clade to develop pelvic fins, the second set of paired fins and the homology (biology), homologous precursor to hindlimbs in tetrapods. 380-million-year-old fossils of three other genera, ''Incisoscutum'', ''M ...
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Superfamily (biology)
In biology, taxonomic rank (which some authors prefer to call nomenclatural rank because ranking is part of nomenclature rather than taxonomy proper, according to some definitions of these terms) is the relative or absolute level of a group of organisms (a ''taxon'') in a hierarchy that reflects evolutionary relationships. Thus, the most inclusive clades (such as Eukarya and Animalia) have the highest ranks, whereas the least inclusive ones (such as ''Homo sapiens'' or '' Bufo bufo'') have the lowest ranks. Ranks can be either relative and be denoted by an indented taxonomy in which the level of indentation reflects the rank, or absolute, in which various terms, such as species, genus, family, order, class, phylum, kingdom, and domain designate rank. This page emphasizes absolute ranks and the rank-based codes (the Zoological Code, the Botanical Code, the Code for Cultivated Plants, the Prokaryotic Code, and thCode for Viruses require them. However, absolute ranks are not r ...
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