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Therizinosaurus
''Therizinosaurus'' (; meaning 'scythe lizard') is a genus of very large therizinosaurid dinosaurs that lived in Asia during the Late Cretaceous period in what is now the Nemegt Formation around 70 million years ago. It contains a single species, ''Therizinosaurus cheloniformis''. The first remains of ''Therizinosaurus'' were found in 1948 by a Mongolian field expedition in the Gobi Desert and later described by Evgeny Maleev in 1954. The genus is only known from a few bones, including gigantic manual unguals (claw bones), from which it gets its name, and additional findings comprising fore and hindlimb elements that were discovered from the 1960s through the 1980s. ''Therizinosaurus'' was a colossal therizinosaurid that could grow up to long and tall, and weigh possibly over . Like other therizinosaurids, it would have been a slow-moving, long-necked, high browser equipped with a rhamphotheca (horny beak) and a wide torso for food processing. Its forelimbs were particularly ...
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Therizinosauridae
Therizinosauridae (meaning 'scythe lizards')Translated paper
is an extinct family of derived (advanced) therizinosauroid dinosaurs whose fossil remains have been found in mostly Late Cretaceous boundary. Even though representative fossils have only been found throughout Asia and North America, the range of Therizinosauridae is believed to have spanned much of the supercontinent of Laurasia based on several footprints and isolated remains in Europe and Africa. As of 2025, the family Therizinosauridae comprises nine definitive genera. Therizinosauridae was named in 1954 by paleontologist Evgeny Maleev after the large, claw-bearing unguals of the type species ''Therizinosaurus cheloniformis''. Therizinosaurids were generally large and very robustly built animals that had a near Convergent evolution, convergent body plan with the mo ...
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Therizinosauria
Therizinosaurs (; once called segnosaurs) are an extinct group of large herbivorous Theropoda, theropod dinosaurs whose fossils have been mainly discovered from Cretaceous deposits in Asia and North America. Potential fragmentary remains have also been found in Jurassic deposits of Asia and Europe. Various features of the forelimbs, skull and pelvis unite these finds as both theropods and maniraptorans, making them relatives of birds. The name of the representative genus, ''Therizinosaurus'', is derived from the Ancient Greek , Greek (, 'to reap' or 'scythe')Translated paper
and (, 'lizard'). The older representative, ''Segnosaurus'', is derived from the Latin ('slow') and the Greek .


History of research


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Evgeny Maleev
Evgeny Aleksandrovich Maleev (, ; 25 February 1915 – 12 April 1966) was a Soviet and Russian paleontologist who did most of his research on reptiles and Asian fossils, such as the naming of the ankylosaur '' Talarurus'' and theropods '' Tarbosaurus'' and '' Therizinosaurus'' along with the family Therizinosauridae. Biography Evgeny Aleksandrovich Maleev was born on February 25, 1915, in the Russian Empire. After being a member on the Eastern Front (World War II) he started his scientific career at the Moscow State University. In 1947 he graduated from the Faculty of Biology, also in that year, he started to work at the Paleontological Institute, Russian Academy of Sciences. In 1950 Maleev presented his PhD thesis, "Morphofunctional analysis of the occipital region of the skull and skeleton of the neck of mammals", being awarded the Candidate of Sciences. Later on, he worked as the deputy manager from 1956 to 1962. He also was the leader of an expedition conducted in 1962 t ...
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1954 In Paleontology
Plants Angiosperms Arthropods Insects Archosauromorphs Phytosaurs Dinosaurs Data courtesy of George Olshevsky's dinosaur genera list. Synapsids Pelycosaurs Eutherians Cetaceans References {{portal, Paleontology 1950s in paleontology Paleontology 4 ...
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Nemegt Formation
The Nemegt Formation (also known as Nemegtskaya Svita) is a geological formation in the Gobi Desert of Mongolia, dating to the Late Cretaceous. The formation consists of river channel sediments and contains fossils of fish, turtles, crocodilians, and a diverse fauna of dinosaurs, including birds. Description The Nemegt Formation is composed of mudstones and sandstones that were deposited by ancient lakes, streams, and flood plains. The Altan Uul locality was described by Michael Novacek as "a canyon carved out of a very rich series of sedimentary rocks" with "steep cliffs and narrow washes". The climate associated with it was wetter than when preceding formations were deposited; there seems to have existed at least some degree of forest cover as indicated by petrified fossil trunks. When examined, the rock facies of the upper section consist of braided meanders, and oxbow lakes. Alteration of sediment grades indicates frequent hydrodynamic condition changes (seasonal rain). Si ...
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Claws
A claw is a curved, pointed appendage found at the end of a toe or finger in most amniotes (mammals, reptiles, birds). Some invertebrates such as beetles and spiders have somewhat similar fine, hooked structures at the end of the leg or tarsus for gripping a surface as they walk. The pincers of crabs, lobsters and scorpions, more formally known as their chelae, are sometimes called claws. A true claw is made of a hard protein called keratin. Claws are used to catch and hold prey in carnivorous mammals such as cats and dogs, but may also be used for such purposes as digging, climbing trees, self-defense and grooming, in those and other species. Similar appendages that are flat and do not come to a sharp point are called nails instead. Claw-like projections that do not form at the end of digits but spring from other parts of the foot are properly named spurs. Tetrapods In tetrapods, claws are made of keratin and consist of two layers. The unguis is the harder external lay ...
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Dinosaur
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutionary history, evolution of dinosaurs is a subject of active research. They became the dominant terrestrial vertebrates after the Triassic–Jurassic extinction event 201.3 mya and their dominance continued throughout the Jurassic and Cretaceous periods. The fossil record shows that birds are feathered dinosaurs, Evolution of birds, having evolved from earlier Theropoda, theropods during the Late Jurassic epoch, and are the only dinosaur lineage known to have survived the Cretaceous–Paleogene extinction event approximately 66 mya. Dinosaurs can therefore be divided into avian dinosaurs—birds—and the extinct non-avian dinosaurs, which are all dinosaurs other than birds. Dinosaurs are varied from taxonomy (biology), taxonomic, ...
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Maniraptora
Maniraptora is a clade of coelurosaurian dinosaurs which includes the birds and the non-avian dinosaurs that were more closely related to them than to ''Ornithomimus velox''. It contains the major subgroups Avialae, Dromaeosauridae, Troodontidae, Oviraptorosauria, and Therizinosauria. ''Ornitholestes'' and the Alvarezsauroidea are also often included. Together with the next closest sister group, the Ornithomimosauria, Maniraptora comprises the more inclusive clade Maniraptoriformes. Maniraptorans first appear in the fossil record during the Jurassic Period (see ''Eshanosaurus''), and origin of birds#Phylogeny, survive today as living birds. Description Maniraptorans are characterized by long arms and three-fingered hands (though reduced or fused in some lineages), as well as a "half-moon shaped" (semilunar bone, semi-lunate) bone in the wrist (Carpal bones, carpus). In 2004, Tom Holtz and Halszka Osmólska pointed out six other maniraptoran characteristics relating to specific de ...
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Ornithischian
Ornithischia () is an extinct clade of mainly herbivorous dinosaurs characterized by a pelvic structure superficially similar to that of birds. The name ''Ornithischia'', or "bird-hipped", reflects this similarity and is derived from the Greek stem ' (), meaning "bird", and ' (), meaning "hip". However, as theropod dinosaurs, birds are only distantly related to this group. Ornithischians with well known anatomical adaptations include the ceratopsians or "horn-faced" dinosaurs (e.g. ''Triceratops''), the pachycephalosaurs or "thick-headed" dinosaurs, the armored dinosaurs (Thyreophora) such as stegosaurs and ankylosaurs, and the ornithopods. There is strong evidence that certain groups of ornithischians lived in herds, often segregated by age group, with juveniles forming their own flocks separate from adults. Some were at least partially covered in filamentous (hair- or feather- like) pelts, and there is much debate over whether these filaments found in specimens of '' Tianyulong ...
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Turtle
Turtles are reptiles of the order (biology), order Testudines, characterized by a special turtle shell, shell developed mainly from their ribs. Modern turtles are divided into two major groups, the Pleurodira (side necked turtles) and Cryptodira (hidden necked turtles), which differ in the way the head retracts. There are 360 living and recently extinct species of turtles, including land-dwelling tortoises and freshwater terrapins. They are found on most continents, some islands and, in the case of sea turtles, much of the ocean. Like other Amniote, amniotes (reptiles, birds, and mammals) they breathe air and do not lay eggs underwater, although many species live in or around water. Turtle shells are made mostly of bone; the upper part is the domed Turtle shell#Carapace, carapace, while the underside is the flatter plastron or belly-plate. Its outer surface is covered in scale (anatomy), scales made of keratin, the material of hair, horns, and claws. The carapace bones deve ...
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Sauropodomorpha
Sauropodomorpha ( ; from Greek, meaning "lizard-footed forms") is an extinct clade of long-necked, herbivorous, saurischian dinosaurs that includes the sauropods and their ancestral relatives. Sauropods generally grew to very large sizes, had long necks and tails, were quadrupedal, and became the largest animals to ever walk the Earth. The primitive sauropodomorphs which preceded the sauropods, and are informally called prosauropods, were smaller and were often able to walk on two legs. The sauropodomorphs were the dominant terrestrial herbivores throughout much of the Mesozoic Era, from their origins in the Late Triassic (approximately 230 Ma) until their decline and extinction at the end of the Cretaceous. History of study Early study Gigantic bones of sauropods have been known for thousands of years and become part of legends and cultures but the beginning of their scientific study was in the 1830s. Most of the early taxonomy was based on incomplete and disarticulated mater ...
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