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Sparassodonta
Sparassodonta (from Ancient Greek, Greek ['], to tear, rend; and , gen. [', '], tooth) is an extinct order (biology), order of carnivore, carnivorous metatherian mammals native to South America, related to modern marsupials. They were once considered to be true marsupials, but are now thought to be a separate side branch that split before the last common ancestor of all modern marsupials. A number of these mammalian predators closely resemble placental predators that evolved separately on other continents, and are cited frequently as examples of convergent evolution. They were first described by Florentino Ameghino, from fossils found in the Santa Cruz beds of Patagonia. Sparassodonts were present throughout South America's long period of "splendid isolation" during the Cenozoic; during this time, they shared the niches for large warm-blooded predators with the flightless Phorusrhacidae, terror birds. Previously, it was thought that these mammals died out in the face of competi ...
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Proborhyaenidae
Proborhyaenidae is an extinct Family (biology), family of metatherian mammals of the order Sparassodonta, which lived in South America from the Eocene (Mustersan) until the Oligocene (Deseadan). Sometimes it has been included as a subfamily of their relatives, the borhyaenids (as Proborhyaeninae). The largest species, ''Proborhyaena gigantea'', is estimated to be about the size of a spectacled bear, with its skull reaching in length, and body mass estimates up to approximately , making the proborhyaenids some of the largest known metatherians. Proborhyaenid remains have been found in western Bolivia, Uruguay, southern Brazil, and the provinces of Mendoza Province, Mendoza, Salta, and Chubut Province, Chubut, in Argentina. Most proborhyaenids had a robust, hyena-like skull, although one species, ''Callistoe vincei'', had an elongate, narrow skull more reminiscent of a thylacine. The teeth were strongly specialized as carnassials for eating meat, and in ''Arminiheringia'' rotated ...
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Borhyaenidae
Borhyaenidae is an extinct metatherian family of low-slung, heavily built predatory mammals in the order Sparassodonta. Borhyaenids are not true marsupials, but members of a sister taxon, Sparassodonta. Like most metatherians, borhyaenids and other sparassodonts are thought to have had a pouch to carry their offspring around. Borhyaenids had strong and powerful jaws, like those of the unrelated placentalians ''Hyaenodon'' and ''Andrewsarchus'', for crushing bones. Borhyaenids grew up to an average of long. Originally, the borhyaenids were considered one of the most diverse groups of sparassodonts, including all species not originally included in the Thylacosmilidae. However, in recent years, with the elevation of most sparassodont subfamilies to family rank and the discovery that borhyaenids are more closely related to proborhyaenids and thylacosmilids than other sparassodonts, the family has been reduced to seven species in four genera. The most studied borhyaenids are the E ...
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Great American Interchange
The Great American Biotic Interchange (commonly abbreviated as GABI), also known as the Great American Interchange and the Great American Faunal Interchange, was an important late Cenozoic paleozoogeographic biotic interchange event in which land and freshwater fauna (animals), fauna migrated from North America to South America via Central America and vice versa, as the volcanic Isthmus of Panama rose up from the sea floor, forming a land bridge between the previously separated continents. Although earlier dispersals had occurred, probably over water, the migration accelerated dramatically about 2.7 million years (Ma (unit), Ma) ago during the Piacenzian age. It resulted from the joining of the Neotropical realm, Neotropic (roughly South American) and Nearctic realm, Nearctic (roughly North American) biogeographic realms definitively to form the Americas. The interchange is visible from observation of both biostratigraphy and nature (neontology). Its most dramatic effect is on the ...
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Metatheria
Metatheria is a mammalian clade that includes all mammals more closely related to marsupials than to placentals. First proposed by Thomas Henry Huxley in 1880, it is a more inclusive group than the marsupials; it contains all marsupials as well as many extinct non-marsupial relatives. It is one of two groups placed in the clade Theria alongside Eutheria, which contains the placentals. Remains of metatherians have been found on all of Earth’s continents. Description Distinctive characteristics ( synapomorphies) of Metatheria include a prehensile tail, the development of a capitular tail on the humerus, the loss of tooth replacement on the second and fifth premolars, lower canines that outwardly diverge from each other, an angular process on the dentary bone—which additionally bears a posterior shelf in its masseteric fossa in Metatheria—that is equal to or greater than the length of the ramus, and the lower fifth premolar with a "very trenchant" cristid obliqua/ecto ...
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Thylacosmilidae
Thylacosmilidae is an extinct family of metatherian predators, related to the modern marsupials, which lived in South America between the Miocene and Pliocene epochs. Like other South American mammalian predators that lived prior to the Great American Biotic Interchange, these animals belonged to the order Sparassodonta, which occupied the ecological niche of many eutherian mammals of the order Carnivora from other continents. The family's most notable feature are the elongated, laterally flattened fangs, which is a remarkable evolutionary convergence with other saber-toothed mammals like '' Barbourofelis'' and ''Smilodon''. Taxonomic history The family Thylacosmilidae was originally erected by Riggs in 1933, to accommodate '' Thylacosmilus'', found in the Pliocene Brochero Formation of Argentina. Later, the family was demoted to a subfamily, as Thylacosmilinae, within Borhyaenidae, a group of superficially canid-like sparassodonts, under the assumption that ''Thylacosmilus ...
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Lycopsis
''Lycopsis'' is an extinct genus of South American metatherian that lived during the Miocene in Argentina and Colombia. History Although not named until 1927, Florentino Ameghino described a species now seen as synonymous with ''Lycopsis torresi'', ''Anatherium oxyrhynchus'', in 1895 based on a mandibular ramus with several teeth.Marshall, L. G. (1977). A new species of Lycopsis (Borhyaenidae: Marsupialia) from the La Venta fauna (Late Miocene) of Colombia, South America. ''Journal of Paleontology'', 633–642. The fossil was recovered from Puesto Estancia La Costa in Santa Cruz, Argentina, dating to the Miocene. The type material of ''Lycopsis'' was collected in July 1895 by "C. Berry" from the middle Miocene strata of the Santa Cruz Formation along the Santa Cruz River in the same area.Cabrera, Á. (1927). Datos para el conocimiento de los dasiuroideos fósiles argentinos. ''Revista del Museo de la Plata'', ''30'', 271–315. The fossils (MLP 11–113) were fragmentary, cons ...
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Patagosmilus
''Patagosmilus'' ("Patagonian knife" in Greek) is an extinct genus of meat-eating metatherian mammal of the family Thylacosmilidae, that lived in the Middle Miocene in South America.''Patagosmilus''
at .org
Like other representatives of this family, such as '' Thylacosmilus atrox'' and '' Anachlysictis gracilis'', it was characterized by its elongated s of the upper jaw, similar to the well know ...
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Phorusrhacidae
Phorusrhacids, colloquially known as terror birds, are an extinct family of large carnivorous, mostly flightless birds that were among the largest apex predators in South America during the Cenozoic era. Their definitive fossil records range from the Middle Eocene to the Late Pleistocene around , though some specimens suggest that they were present since the Early Eocene. They ranged in height from . One of the largest specimens from the Early Pleistocene of Uruguay, possibly belonging to '' Devincenzia'', would have weighed up to . Their closest modern-day relatives are believed to be the seriemas. '' Titanis walleri'', one of the larger species, is known from Texas and Florida in North America. This makes the phorusrhacids the only known large South American predator to migrate north in the Great American Interchange that followed the formation of the Isthmus of Panama land bridge (the main pulse of the interchange began about 2.6 Ma ago; ''Titanis'' at 5 Ma was an early northwa ...
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Carnivore
A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant Plants are the eukaryotes that form the Kingdom (biology), kingdom Plantae; they are predominantly Photosynthesis, photosynthetic. This means that they obtain their energy from sunlight, using chloroplasts derived from endosymbiosis with c ... whose nutrition and energy requirements are met by eating, consumption of animal tissue (biology), tissues (mainly muscle, adipose tissue, fat and other soft tissues) as food, whether through predation or scavenger, scavenging. Nomenclature Mammal order The technical term for mammals in the order (biology), order Carnivora is ''carnivoran'', and they are so-named because most member species in the group have a carnivorous diet, but the similarity of the name of the order and the name of the diet causes confusion. Many but not all carnivorans are meat eaters; a few, such as the fe ...
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Pliocene
The Pliocene ( ; also Pleiocene) is the epoch (geology), epoch in the geologic time scale that extends from 5.33 to 2.58See the 2014 version of the ICS geologic time scale
million years ago (Ma). It is the second and most recent epoch of the Neogene Period in the Cenozoic, Cenozoic Era. The Pliocene follows the Miocene Epoch and is followed by the Pleistocene Epoch. Prior to the 2009 revision of the geologic time scale, which placed the four most recent major glaciations entirely within the Pleistocene, the Pliocene also included the Gelasian Stage, which lasted from 2.59 to 1.81 Ma, and is now included in the Pleistocene. As with other older geologic periods, the Stratum, geological strata that define the start and end are well-identified but the exact dates of the start a ...
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Hondadelphidae
''Hondadelphys'' is an extinct genus of carnivorous sparassodonts, known from the Middle Miocene of Colombia. The type species, ''H. fieldsi'', was described in 1976 from the fossil locality of La Venta, which hosts fossils from the Villavieja Formation. ''Hondadelphys ''was originally interpreted as belonging to the opossum family Didelphidae, but subsequently assigned to its own family, Hondadelphidae and interpreted as a basal sparassodont. The genus name refers to the Honda Group, the stratigraphic group in which the fossils of this animal were first found, combined with ''delphys ''(Greek for "womb", a common suffix used for opossum-like metatherian Metatheria is a mammalian clade that includes all mammals more closely related to marsupials than to placentals. First proposed by Thomas Henry Huxley in 1880, it is a more inclusive group than the marsupials; it contains all marsupials as well ...s). References Sparassodonta Miocene mammals of South America L ...
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