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Skoliomonas
''Skoliomonas'' is a genus of anaerobic protists closely related to barthelonids, a small group of basal eukaryotes within the phylum Metamonada. It is a monotypic genus containing the sole species ''Skoliomonas litria''. Members of this genus are informally named skoliomonads. They are found inhabiting hypersaline alkaline lakes in Tanzania and North America. Etymology The name ''Skoliomonas'' derives , 'crooked', which refers to the hunched appearance of the cell and the twisted venral groove, and ''μονάς (monas)'' 'unit', a common suffix used for unicellular protists. The specific epithet ''litria'' comes , an alternative form of ''νίτρον (nitron)'', meaning sodium carbonate; the term was used by Herodotus to describe embalming salts used for mummification in Ancient Egypt, harvested from carbonate-rich soda lakes of the Natron Valley. The authors of the species chose the African-specific version of the Ancient Greek word to reflect the African type locality and ...
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Metamonad
The metamonads are a large group of flagellate amitochondriate microscopic eukaryotes. They include the retortamonads, diplomonads, parabasalids, oxymonads, and a range of more poorly studied taxa, most of which are free-living flagellates. All metamonads are anaerobic (many being aerotolerant anaerobes), and most members of the four groups listed above are symbiotes or parasites of animals, as is the case with ''Giardia lamblia'' which causes diarrhea in mammals. Characteristics A number of parabasalids and oxymonads are found in termite guts, and play an important role in breaking down the cellulose found in wood. Some other metamonads are parasites. These flagellates are unusual in lacking aerobic mitochondria. Originally they were considered among the most primitive eukaryotes, diverging from the others before mitochondria appeared. However, they are now known to have lost aerobic mitochondria secondarily, and retain both organelles and nuclear genes derived ultimate ...
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Protist
A protist ( ) or protoctist is any eukaryotic organism that is not an animal, land plant, or fungus. Protists do not form a natural group, or clade, but are a paraphyletic grouping of all descendants of the last eukaryotic common ancestor excluding land plants, animals, and fungi. Protists were historically regarded as a separate taxonomic kingdom known as Protista or Protoctista. With the advent of phylogenetic analysis and electron microscopy studies, the use of Protista as a formal taxon was gradually abandoned. In modern classifications, protists are spread across several eukaryotic clades called supergroups, such as Archaeplastida ( photoautotrophs that includes land plants), SAR, Obazoa (which includes fungi and animals), Amoebozoa and " Excavata". Protists represent an extremely large genetic and ecological diversity in all environments, including extreme habitats. Their diversity, larger than for all other eukaryotes, has only been discovered in rece ...
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Barthelonid
''Barthelona'' is a genus of anaerobic protists. They are basal eukaryotes closely related to skoliomonads, within the phylum Metamonada. It is a monotypic genus containing the sole species ''Barthelona vulgaris''. Members of this genus are informally known as barthelonids. Description Barthelonids are flagellates, unicellular protists (a type of eukaryote) that use flagella for movement. Their cells measure 6–8 μm in length, and are oval on the dorsal side and flattened on the ventral side. Each cell has two flagella inserted sub-apically and laterally, at an angle of 60° from each other. The anterior flagellum is 1–2 times the cell length, while the posterior flagellum is 3–5 times as large as the cell body, and is slightly acronematic. The nucleus is located in the anterior end of the cell. The cytoplasm usually contains refractile granules. Barthelonids are free-living solitary heterotrophs. They swim or, more rarely, glide slowly. When stationary, they still be ...
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Genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In binomial nomenclature, the genus name forms the first part of the binomial species name for each species within the genus. :E.g. ''Panthera leo'' (lion) and ''Panthera onca'' (jaguar) are two species within the genus ''Panthera''. ''Panthera'' is a genus within the family Felidae. The composition of a genus is determined by taxonomy (biology), taxonomists. The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including the idea that a newly defined genus should fulfill these three criteria to be descriptively useful: # monophyly – all descendants of an ancestral taxon are grouped together (i.e. Phylogeneti ...
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Cytoplasm
The cytoplasm describes all the material within a eukaryotic or prokaryotic cell, enclosed by the cell membrane, including the organelles and excluding the nucleus in eukaryotic cells. The material inside the nucleus of a eukaryotic cell and contained within the nuclear membrane is termed the nucleoplasm. The main components of the cytoplasm are the cytosol (a gel-like substance), the cell's internal sub-structures, and various cytoplasmic inclusions. In eukaryotes the cytoplasm also includes the nucleus, and other membrane-bound organelles.The cytoplasm is about 80% water and is usually colorless. The submicroscopic ground cell substance, or cytoplasmic matrix, that remains after the exclusion of the cell organelles and particles is groundplasm. It is the hyaloplasm of light microscopy, a highly complex, polyphasic system in which all resolvable cytoplasmic elements are suspended, including the larger organelles such as the ribosomes, mitochondria, plant plasti ...
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Fornicata
Trichozoa is a group of excavates. "Fornicata" is a similar grouping, but it excludes Parabasalia.Tree
at National Institute of Genetics
"Eopharyngia" is an even more narrow grouping, including Retortamonadida and Diplomonadida but not Carpediemonas.


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Soda Lake
A soda lake or alkaline lake is a lake on the strongly base (chemistry), basic side of neutrality, typically with a pH, pH value between 9 and 12. They are characterized by high concentrations of carbonate salts, typically sodium carbonate (and related salt complexes), giving rise to their alkalinity. In addition, many soda lakes also contain high concentrations of sodium chloride and other dissolved Salt (chemistry), salts, making them Salt lake, saline or Hypersaline lake, hypersaline lakes as well. High pH and salinity often coincide, because of how soda lakes develop. The resulting hypersaline and highly alkalic soda lakes are considered some of the most extreme aquatic environments on Earth. In spite of their apparent inhospitability, soda lakes are often highly Productivity (ecology), productive ecosystems, compared to their (pH-neutral) freshwater counterparts. Primary production#GPP and NPP, Gross primary production (photosynthesis) rates above (grams of carbon per squar ...
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Anaerobe
An anaerobic organism or anaerobe is any organism that does not require molecular oxygen for growth. It may react negatively or even die if free oxygen is present. In contrast, an aerobic organism (aerobe) is an organism that requires an oxygenated environment. Anaerobes may be unicellular (e.g. protozoans, bacteria) or multicellular. Most fungi are obligate aerobes, requiring oxygen to survive. However, some species, such as the Chytridiomycota that reside in the rumen of cattle, are obligate anaerobes; for these species, anaerobic respiration is used because oxygen will disrupt their metabolism or kill them. The sea floor is possibly one of the largest accumulation of anaerobic organisms on Earth, where microbes are primarily concentrated around hydrothermal vents. These microbes produce energy in absence of sunlight or oxygen through a process called chemosynthesis, whereby inorganic compounds such as hydrogen gas, hydrogen sulfide or ferrous ions are converted into organic m ...
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Carpediemonas
''Carpediemonas'' is genus of Metamonada, and belongs to the group Excavata. This organism is a unicellular flagellated eukaryote that was first discovered in substrate samples from the Great Barrier Reef. ''Carpediemonas'' can be found in anaerobic intertidal sediment, where it feeds on bacteria. A feature of this species is the presence of a feeding groove, a characteristic of the excavates. Like most other metamonads, ''Carpediemonas'' does not rely on an aerobic mitochondrion to produce energy. Instead, it contains hydrogenosomes that are used to produce ATP. This organism has two flagella: a posterior one used for feeding on the substrate, and an anterior one that moves in a slower sweeping motion. ''Carpediemonas'' is assigned to the fornicates, where similar ''Carpediemonas''-like organisms are used in researching the evolution within excavates. Although ''Carpediemonas'' is a member of the metamonads, it is unusual in the sense that it is free-living and has three basal bo ...
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Cytopharynx
A cytostome (from ''cyto-'', cell and ''stome-'', mouth) or cell mouth is a part of a cell specialized for phagocytosis, usually in the form of a microtubule-supported funnel or groove. Food is directed into the cytostome, and sealed into vacuoles. Only certain groups of protozoa, such as the Ciliophora and Excavata, have cytostomes. An example is ''Balantidium coli'', a ciliate. In other protozoa, and in cells from multicellular organisms, phagocytosis takes place at any point on the cell or feeding takes place by absorption. Structure The cytostome forms an invagination on the cell surface and is typically directed towards the nucleus of the cell.Okuda, Kendi, et al. "The cytostome of Trypanosoma cruzi epimastigotes is associated with the flagellar complex." Experimental parasitology 92.4 (1999): 223-231. The cytostome is often labeled as the entire invagination, but in fact the cytostome only constitutes the opening of the invagination at the surface of the cell. The rest o ...
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Microbial Cyst
A microbial cyst is a resting or dormant stage of a microorganism, that can be thought of as a state of suspended animation in which the metabolic processes of the cell are slowed and the cell ceases all activities like feeding and locomotion. Many groups of single-celled, microscopic organisms, or microbes, possess the ability to enter this dormant state. Encystment, the process of cyst formation, can function as a method for dispersal and as a way for an organism to survive in unfavorable environmental conditions. These two functions can be combined when a microbe needs to be able to survive harsh conditions between habitable environments (such as between hosts) in order to disperse. Cysts can also be sites for nuclear reorganization and cell division, and in parasitic species they are often the infectious stage between hosts. When the encysted microbe reaches an environment favorable to its growth and survival, the cyst wall breaks down by a process known as excystation. E ...
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