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Sedentaria
Sedentaria is a diverse clade of annelid worms. It is traditionally treated as a subclass of the paraphyletic class Polychaeta, but it is also a monophyletic group uniting several polychaetes and the monophyletic class Clitellata. It is the sister group of Errantia. Sedentaria are mainly found within marine environments that have low oxygen levels  and are specially adapted to these low oxygen environments by  increasing gill surface area and having high-affinity respiratory proteins. Furthermore, they go through a process of metabolic depression which lowers their energy use so that they can inhibit these low oxygen zones . Phylogeny The phylogeny of polychaetes is slowly being resolved. Sedentaria and Errantia are the two biggest clades of polychaetes, and together they compose clade Pleistoannelida. Sedentaria's most basal clade is Orbiniida. Other groups that are nested within Sedentaria are: Clitellata, the Sabellida/ Spionida clade, Opheliida, Echiura, Cirratulifor ...
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Pleistoannelida
Pleistoannelida is a group of annelid worms that comprises the vast majority of the diversity in phylum Annelida. Discovered through phylogenetic analyses, it is the largest clade of annelids, comprised by the last common ancestor of the highly diverse sister groups Errantia and Sedentaria (Clitellata and related polychaetes) and all the descendants of that ancestor. Most groups in the clade find their ancestors within the Cambrian explosion when Annelid diversity expanded dramatically. The Pleistoannelida clade covers a variety of traits. However, the evolution of simple to complex eyes, developed papillae for burrowing, and for some specialized radioles for feeding can be seen universally across every species. New findings have discovered the range of Annelid diversity have led to uncertainty if groups with developed ancestral traits should remain within the clade. Furthermore, there's been a lack of recently discovered Annelid traits being used in the categorization of groups wit ...
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Errantia
Errantia is a diverse group of marine life, marine polychaete worms in the phylum Annelida. Traditionally a subclass (biology), subclass of the paraphyletic class Polychaeta, it is currently regarded as a monophyletic group within the larger Pleistoannelida, composed of Errantia and Sedentaria. These worms are found worldwide in marine environments and brackish water. Phylogeny The phylogeny of polychaetes is slowly being resolved. Errantia and Sedentaria are the two biggest clades of polychaetes, and together they compose clade Pleistoannelida. Two groups are nested within Errantia: Aciculata (Eunicida + Phyllodocida) and Protodriliformia (progenesis, small meiofaunal worms such as the Protodrilida). Historically, the order Amphinomida was part of this subclass. However, phylogenetic analyses place Amphinomida inside a basal clade with Sipunculida and ''Lobatocerebrum'', and this clade is the sister group to Pleistoannelida. Some taxon, taxa, such as Spintheridae and Myzostomida ...
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Orbiniida
Orbiniida is an order of small polychaete worms in the phylum Annelida. It is the earliest diverging clade in Sedentaria. Along with Protodriliformia (in Errantia), this order is composed of meiofaunal marine worms formerly known as " archiannelids". These worms inhabit the marine interstitial ecosystem, the space between sand grains. Classification First mentioned in 1977, Orbiniida was an order of the polyphyletic infraclass " Scolecida" of the class " Polychaeta". It was composed of the families Orbiniidae, Paraonidae and Questidae, all of which share in common the lack of antennae and palps, and the presence of an eversible pharynx and biramous parapodia with simple chaetae. However, through phylogenetic analysis it was found that Questidae and Paraonidae are more closely related to Clitellata and Cirratulida respectively. Because of this, the taxonomic composition of Orbiniida was changed to exclude these two families and include several interstitial groups pr ...
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Annelid
The annelids (), also known as the segmented worms, are animals that comprise the phylum Annelida (; ). The phylum contains over 22,000 extant species, including ragworms, earthworms, and leeches. The species exist in and have adapted to various ecologies – some in marine environments as distinct as tidal zones and hydrothermal vents, others in fresh water, and yet others in moist terrestrial environments. The annelids are bilaterally symmetrical, triploblastic, coelomate, invertebrate organisms. They also have parapodia for locomotion. Most textbooks still use the traditional division into polychaetes (almost all marine), oligochaetes (which include earthworms) and leech-like species. Cladistic research since 1997 has radically changed this scheme, viewing leeches as a sub-group of oligochaetes and oligochaetes as a sub-group of polychaetes. In addition, the Pogonophora, Echiura and Sipuncula, previously regarded as separate phyla, are now regarded as sub-grou ...
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Polychaetes
Polychaeta () is a paraphyletic class of generally marine annelid worms, commonly called bristle worms or polychaetes (). Each body segment has a pair of fleshy protrusions called parapodia that bear many bristles, called chaetae, which are made of chitin. More than 10,000 species are described in this class. Common representatives include the lugworm (''Arenicola marina'') and the sandworm or clam worm ''Alitta''. Polychaetes as a class are robust and widespread, with species that live in the coldest ocean temperatures of the abyssal plain, to forms which tolerate the extremely high temperatures near hydrothermal vents. Polychaetes occur throughout the Earth's oceans at all depths, from forms that live as plankton near the surface, to a 2- to 3-cm specimen (still unclassified) observed by the robot ocean probe ''Nereus'' at the bottom of the Challenger Deep, the deepest known spot in the Earth's oceans. Only 168 species (less than 2% of all polychaetes) are known from fr ...
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Polychaete
Polychaeta () is a paraphyletic class of generally marine Annelid, annelid worms, common name, commonly called bristle worms or polychaetes (). Each body segment has a pair of fleshy protrusions called parapodia that bear many bristles, called chaetae, which are made of chitin. More than 10,000 species are described in this class. Common representatives include the lugworm (''Arenicola marina'') and the Alitta virens, sandworm or Alitta succinea, clam worm ''Alitta''. Polychaetes as a class are robust and widespread, with species that live in the coldest ocean temperatures of the abyssal plain, to forms which tolerate the extremely high temperatures near hydrothermal vents. Polychaetes occur throughout the Earth's oceans at all depths, from forms that live as plankton near the surface, to a 2- to 3-cm specimen (still unclassified) observed by the robot ocean probe Nereus (underwater vehicle), ''Nereus'' at the bottom of the Challenger Deep, the deepest known spot in the Earth's ...
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Arenicolidae
Arenicolidae is a family of marine polychaete worms. They are commonly known as lugworms and the little coils of sand they produce are commonly seen on the beach. Arenicolids are found worldwide, mostly living in burrows in sandy substrates. Most are detritivores but some graze on algae. Description The arenicolids are characterised by an elongated cylindrical body separated into two or three distinct regions. The prostomium has no appendages or palps. There are one or two anterior segments without setae. On the other segments, all the setae are unbranched, including the capillary setae and the rostrate uncini. The notopodia are bluntly truncate and the neuropodia are elongated tori forming long transverse welts in some of the setigers. The notosetae have either a capillary function or act as limbs and the neurosetae are rostrate hooks. There are branchiae present on some of the setigers in the middle or posterior regions. Apart from the genus '' Branchiomaldane'', the lugwo ...
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Maldanidae
Maldanidae is a family of more than 200 species of marine polychaetes commonly known as bamboo worms or maldanid worms. They belong to the order Capitellida, in the phylum Annelida. They are most closely related to family Arenicolidae, and together form the clade Maldanomorpha. Morphology Maldanid worms have a long and cylindrical body that usually bends at one or both ends. It can be divided into four parts: 1) a head, formed by a dorsally positioned prostomium that is fused to the peristomium, sometimes with a flattened cephalic plate; 2) a thorax, formed by the first four chaetigers (i.e. chaetae-bearing segments), usually with strong spines; 3) an abdomen, with several longer chaetigers that are often followed by a number of achaetous (i.e. without chaetae) segments; 4) a posterior end, with a pygidium that contains the anus. The maldanid pygidium takes a wide variety of forms such as conical, plate-shaped and funnel-shape. Sometimes a few prepygidial (i.e. anterior to the ...
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Spintheridae
Spintheridae is a family of marine polychaete worms with a single genus, ''Spinther'', containing these species: * '' Spinther alaskensis'' Hartman, 1948 * '' Spinther arcticus'' (M. Sars, 1851) (includes '' Spinther miniaceus'' Grube, 1860) *'' Spinther australiensis'' Augener, 1913 * '' Spinther citrinus'' (Stimpson, 1854) *'' Spinther ericinus'' Yamamoto & Imajima, 1985 *'' Spinther hystrix'' Uschakov, 1950 * '' Spinther japonicus'' Imajima and Hartman, 1964 * '' Spinther oniscoides'' Johnston, 1845 *'' Spinther sagamiensis'' Imajima, 2003 *'' Spinther usarpia'' Hartman, 1967 * '' Spinther vegae'' Augener, 1928 (includes '' Spinther wireni'' Hartman, 1948) The animal lives as a symbiont on sponges. Johnston's paper does not explain the choice of the name, but ancient Greek σπινθήρ means "spark." In images of the living animal, it appears to be surrounded by a cloud of pinpoints of light. References Polychaetes Taxa named by George Johnston (naturalist) {{an ...
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Taxon
In biology, a taxon (back-formation from ''taxonomy''; : taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and given a particular ranking, especially if and when it is accepted or becomes established. It is very common, however, for taxonomists to remain at odds over what belongs to a taxon and the criteria used for inclusion, especially in the context of rank-based (" Linnaean") nomenclature (much less so under phylogenetic nomenclature). If a taxon is given a formal scientific name, its use is then governed by one of the nomenclature codes specifying which scientific name is correct for a particular grouping. Initial attempts at classifying and ordering organisms (plants and animals) were presumably set forth in prehistoric times by hunter-gatherers, as suggested by the fairly sophisticated folk taxonomies. Much later, Aristotle, and later st ...
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Monophyletic
In biological cladistics for the classification of organisms, monophyly is the condition of a taxonomic grouping being a clade – that is, a grouping of organisms which meets these criteria: # the grouping contains its own most recent common ancestor (or more precisely an ancestral population), i.e. excludes non-descendants of that common ancestor # the grouping contains all the descendants of that common ancestor, without exception Monophyly is contrasted with paraphyly and polyphyly as shown in the second diagram. A ''paraphyletic'' grouping meets 1. but not 2., thus consisting of the descendants of a common ancestor, excepting one or more monophyletic subgroups. A '' polyphyletic'' grouping meets neither criterion, and instead serves to characterize convergent relationships of biological features rather than genetic relationships – for example, night-active primates, fruit trees, or aquatic insects. As such, these characteristic features of a polyphyletic grouping ...
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