Scleropyrenium Kurokawae
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Scleropyrenium Kurokawae
''Scleropyrenium'' is a genus of squamulose (scaly) lichens in the family Verrucariaceae. It has two species. The genus was circumscribed in 1993 by Japanese lichenologist Hiroshi Harada, with '' S. japonicum'' as the type species. Characteristics of the genus include a dark brown to almost black exciple (the rim of tissue around the hymenium), pycnidia of the '' Staurothele''-type, and a pachydermatous upper cortex Cortex or cortical may refer to: Biology * Cortex (anatomy), the outermost layer of an organ ** Cerebral cortex, the outer layer of the vertebrate cerebrum, part of which is the ''forebrain'' *** Motor cortex, the regions of the cerebral cortex i .... References Verrucariales Lichen genera Eurotiomycetes genera Taxa described in 1993 {{Eurotiomycetes-stub ...
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Genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In binomial nomenclature, the genus name forms the first part of the binomial species name for each species within the genus. :E.g. ''Panthera leo'' (lion) and ''Panthera onca'' (jaguar) are two species within the genus ''Panthera''. ''Panthera'' is a genus within the family Felidae. The composition of a genus is determined by taxonomy (biology), taxonomists. The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including the idea that a newly defined genus should fulfill these three criteria to be descriptively useful: # monophyly – all descendants of an ancestral taxon are grouped together (i.e. Phylogeneti ...
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Squamulose Lichen
A squamulose lichen is a lichen that is composed of small, often overlapping "scales" called . If they are raised from the substrate and appear leafy, the lichen may appear to be a foliose lichen, but the underside does not have a "skin" (cortex), as foliose lichens do. Squamulose lichens are composed of flattish units that are usually tightly clustered. They are like an intermediate between crustose Crustose is a Habit (biology), habit of some types of algae and lichens in which the organism grows tightly appressed to a substrate, forming a biological layer. ''Crustose'' adheres very closely to the Substrate (biology), substrates at all poin ... and foliose lichens. Examples of squamulose lichens include '' Vahliella leucophaea'', '' Cladonia subcervicornis'' and '' Lichenomphalia hudsoniana''. References Lichenology {{lichen-stub ...
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Lichen
A lichen ( , ) is a hybrid colony (biology), colony of algae or cyanobacteria living symbiotically among hypha, filaments of multiple fungus species, along with yeasts and bacteria embedded in the cortex or "skin", in a mutualism (biology), mutualistic relationship.Introduction to Lichens – An Alliance between Kingdoms
. University of California Museum of Paleontology. .
Lichens are the lifeform that first brought the term symbiosis (as ''Symbiotismus'') into biological context. Lichens have since been recognized as important actors in nutrient cycling and producers which many higher trophic feeders feed on, such as reindeer, gastropods, nematodes, mites, and springtails. Lichens have properties different from those of their component organisms. They come in man ...
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Verrucariaceae
Verrucariaceae is a family of lichens and a few non-lichenised fungi in the order Verrucariales. The lichens have a wide variety of thallus forms, from crustose (crust-like) to foliose (bushy) and squamulose (scaly). Most of them grow on land, some in freshwater and a few in the sea. Many are free-living but there are some species that are parasites on other lichens, while one marine species always lives together with a leafy green alga. Several characteristics of the spore-bearing structures, the ascomata, define the family, including their perithecioid form–more or less spherical or flask-shaped, with a single opening and otherwise completely enclosed by a wall. Squamulose members of the Verrucariaceae with simple ascospores (lacking partitions called septa), and without algae in the spore-bearing region are known as lichens; there are more than 80 of these species. The family has several dozen lichenicolous (lichen-dwelling) examples, including a few genera that co ...
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Circumscription (taxonomy)
In biological taxonomy, circumscription is the content of a taxon, that is, the delimitation of which subordinate taxa are parts of that taxon. For example, if we determine that species X, Y, and Z belong in genus A, and species T, U, V, and W belong in genus B, those are our circumscriptions of those two genera. Another systematist might determine that T, U, V, W, X, Y, and Z all belong in genus A. Agreement on circumscriptions is not governed by the Codes of Zoological or Botanical Nomenclature, and must be reached by scientific consensus. A goal of biological taxonomy is to achieve a stable circumscription for every taxon. This goal conflicts, at times, with the goal of achieving a natural classification that reflects the evolutionary history of divergence of groups of organisms. Balancing these two goals is a work in progress, and the circumscriptions of many taxa that had been regarded as stable for decades are in upheaval in the light of rapid developments in molecu ...
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Type Species
In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological Type (biology), type wiktionary:en:specimen, specimen (or specimens). Article 67.1 A similar concept is used for suprageneric groups and called a type genus. In botanical nomenclature, these terms have no formal standing under the International Code of Nomenclature for algae, fungi, and plants, code of nomenclature, but are sometimes borrowed from zoological nomenclature. In botany, the type of a genus name is a specimen (or, rarely, an illustration) which is also the type of a species name. The species name with that type can also be referred to as the type of the genus name. Names of genus and family ranks, the various subdivisions of those ranks, and some higher-rank names based on genus names, have suc ...
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Hymenium
The hymenium is the tissue layer on the hymenophore of a fungal fruiting body where the cells develop into basidia or asci, which produce spores. In some species all of the cells of the hymenium develop into basidia or asci, while in others some cells develop into sterile cells called cystidia ( basidiomycetes) or paraphyses ( ascomycetes). Cystidia are often important for microscopic identification. The subhymenium consists of the supportive hyphae from which the cells of the hymenium grow, beneath which is the hymenophoral trama, the hyphae that make up the mass of the hymenophore. The position of the hymenium is traditionally the first characteristic used in the classification and identification of mushrooms. Below are some examples of the diverse types which exist among the macroscopic Basidiomycota and Ascomycota. * In agarics, the hymenium is on the vertical faces of the gills. * In boletes and polypores, it is in a spongy mass of downward-pointing tubes ...
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Pycnidia
A pycnidium (plural pycnidia) is an asexual fruiting body produced by mitosporic fungi, for instance in the order Sphaeropsidales ( Deuteromycota, Coelomycetes) or order Pleosporales (Ascomycota, Dothideomycetes). It is often spherical or inversely pearshaped ( obpyriform) and its internal cavity is lined with conidiophore A conidium ( ; : conidia), sometimes termed an asexual chlamydospore or chlamydoconidium (: chlamydoconidia), is an Asexual reproduction, asexual, non-motility, motile spore of a fungus. The word ''conidium'' comes from the Ancient Greek word f ...s. When ripe, an opening generally appears at the top, through which the pycnidiospores escape. References {{reflist Further reading *Kulik, Martin M. "Symptomless infection, persistence, and production of pycnidia in host and non-host plants by Phomopsis batatae, Phomopsis phaseoli, and Phomopsis sojae, and the taxonomic implications." Mycologia(1984): 274–291. *Calpouzos, L., and D. B. Lapis. "Effects of l ...
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Staurothele
''Staurothele'' is a genus of saxicolous (rock-dwelling), crustose lichens in the family Verrucariaceae. It has about 40 species. When the fungus is part of a lichen A lichen ( , ) is a hybrid colony (biology), colony of algae or cyanobacteria living symbiotically among hypha, filaments of multiple fungus species, along with yeasts and bacteria embedded in the cortex or "skin", in a mutualism (biology), m ..., the genus of lichen is commonly called rock pimples.Field Guide to California Lichens, Stephen Sharnoff, Yale University Press, 2014, Species *'' Staurothele alboterrestris'' *'' Staurothele ambrosiana'' *'' Staurothele arctica'' *'' Staurothele areolata'' *'' Staurothele bacilligera'' *'' Staurothele caesia'' *'' Staurothele clopima'' *'' Staurothele dendritica'' *'' Staurothele desquamescens'' *'' Staurothele drummondii'' *'' Staurothele elenkinii'' *'' Staurothele fissa'' *'' Staurothele frustulenta'' *'' Staurothele geoica'' *'' Staurothele guestp ...
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Cortex (botany)
In botany, a cortex is an outer layer of a stem or root in a vascular plant, lying below the epidermis but outside of the vascular bundles. The cortex is composed mostly of large thin-walled parenchyma cells of the ground tissue system and shows little to no structural differentiation. The outer cortical cells often acquire irregularly thickened cell walls, and are called collenchyma cells. Plants Stems and branches In the three dimensional structure of herbaceous stems, the epidermis, cortex and vascular cambium form concentric cylinders around the inner cylindrical core of pith. Some of the outer cortical cells may contain chloroplasts, giving them a green color. They can therefore produce simple carbohydrates through photosynthesis. In woody plants, the cortex is located between the periderm (bark) and the vascular tissue ( phloem, in particular). It is responsible for the transportation of materials into the central cylinder of the root through diffusion and ma ...
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