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Rafetus Bohemicus
''Rafetus bohemicus'' is an extinct species of softshell turtle from the Early Miocene (Burdigalian) of the Czech Republic. Specimens have been found at the northeastern margin of the Most Basin at a fossil site called Břešt’any. Fossil carapaces of "'' R. pontanus''", a potential taxonomic synonym (although currently considered a ''nomen dubium''), have been found in slightly younger rocks in Lom and Louka u Litvínova. Geology The composition of the Břešt’any Clay is primarily limnic ceramic clay, with the occasional siderite concretion. It has been dated to ~17.5 Ma. It is located between the layers of rock that make up the Holešice and Libkovice Formations. This fine, kaolinite rich mudstone preserves fossils from an Early Miocene (Burdigalian) lacustrine environment, during a period of global warming known as the Mid-Miocene Climatic Optimum. Here, fossils of ''Rafetus bohemicus'' are preserved in either limnic clays or siderite concretions; those of the former ...
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Trionychidae
Trionychidae is a family of turtles, commonly known as softshell turtles or simply softshells. The family was described by Leopold Fitzinger in 1826. Softshells include some of the world's largest freshwater turtles, though many can adapt to living in highly brackish waters. Members of this family occur in Africa, Asia, and North America, with extinct species known from Australia. Most species have traditionally been included in the genus '' Trionyx'', but the vast majority have since been moved to other genera. Among these are the North American '' Apalone'' softshells that were placed in ''Trionyx'' until 1987. Characteristics Turtles of the family Trionychidae are called "softshell" because their carapaces lack horny scutes (scales), though the spiny softshell, '' Apalone spinifera'', does have some scale-like projections, to which its common name refers. The carapace is leathery and pliable, particularly at the sides. The central part of the carapace has a layer of solid ...
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The Reconstruction Of Skulls Of Rafetus Species
''The'' is a grammatical article in English, denoting nouns that are already or about to be mentioned, under discussion, implied or otherwise presumed familiar to listeners, readers, or speakers. It is the definite article in English. ''The'' is the most frequently used word in the English language; studies and analyses of texts have found it to account for seven percent of all printed English-language words. It is derived from gendered articles in Old English which combined in Middle English and now has a single form used with nouns of any gender. The word can be used with both singular and plural nouns, and with a noun that starts with any letter. This is different from many other languages, which have different forms of the definite article for different genders or numbers. Pronunciation In most dialects, "the" is pronounced as (with the voiced dental fricative followed by a schwa) when followed by a consonant sound, and as (homophone of the archaic pronoun ''thee'' ...
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Miocene Turtles
The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recent" because it has 18% fewer modern marine invertebrates than the Pliocene has. The Miocene followed the Oligocene and preceded the Pliocene. As Earth went from the Oligocene through the Miocene and into the Pliocene, the climate slowly cooled towards a series of ice ages. The Miocene boundaries are not marked by distinct global events but by regionally defined transitions from the warmer Oligocene to the cooler Pliocene Epoch. During the Early Miocene, Afro-Arabia collided with Eurasia, severing the connection between the Mediterranean and Indian Oceans, and allowing the interchange of fauna between Eurasia and Africa, including the dispersal of proboscideans and hominoids into Eurasia. During the late Miocene, the connections between the ...
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Burdigalian Life
The Burdigalian is, in the geologic timescale, an age (geology), age or stage (stratigraphy), stage in the early Miocene. It spans the time between 20.43 ± 0.05 annum, Ma and 15.97 ± 0.05 Ma (million years ago). Preceded by the Aquitanian (stage), Aquitanian, the Burdigalian was the first and longest warming period of the MioceneEdward Petuch, Ph.D. Florida Atlantic University, Department of Geosciences. and is succeeded by the Langhian. Stratigraphic definition The name Burdigalian comes from ''Burdigala'', the Latin name for the city of Bordeaux, France. The Burdigalian Stage was introduced in scientific literature by Charles Depéret in 1892. The base of the Burdigalian is at the first appearance of foram species ''Globigerinoides altiaperturus'' and the top of magnetic chronozone C6An. , an official GSSP for the Burdigalian had not yet been assigned. The top of the Burdigalian (the base of the Langhian) is defined by the first appearance of foram species ''Praeorbulina gl ...
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Fossil Taxa Described In 1930
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in amber, hair, petrified wood and DNA remnants. The totality of fossils is known as the ''fossil record''. Though the fossil record is incomplete, numerous studies have demonstrated that there is enough information available to give a good understanding of the pattern of diversification of life on Earth. In addition, the record can predict and fill gaps such as the discovery of ''Tiktaalik'' in the arctic of Canada. Paleontology includes the study of fossils: their age, method of formation, and evolutionary significance. Specimens are sometimes considered to be fossils if they are over 10,000 years old. The oldest fossils are around 3.48 billion years to 4.1 billion years old. Early edition, published online before print. The ob ...
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Rafetus
''Rafetus'' is a genus of highly endangered softshell turtles in the family Trionychidae. It is a genus of large turtles which are found in freshwater habitats in Eurasia. Taxonomy Evolution Phylogenetic evidence largely suggests that ''Rafetus'' is most closely related to the widespread North American genus ''Apalone''. The two genera appear to have diverged during the Late Eocene, likely when the ancestors of ''Apalone'' dispersed into North America via Beringia. Species According to most taxonomists, the genus ''Rafetus'' contains the following two extant species. *''Rafetus euphraticus'' – Euphrates softshell turtle; Tigris and Euphrates Rivers in Iraq, Syria, Turkey, and Khūzestān Province of Iran. IUCN: Endangered. *''Rafetus swinhoei'' – Yangtze giant softshell turtle; Only three known living individuals, one in Suzhou Zoo (China) and two in two lakes in northern Vietnam, Dong Mo lake and Xuan Khanh lake. IUCN: Critically endangered. A possible third sp ...
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Euphrates Softshell Turtle
The Euphrates softshell turtle (''Rafetus euphraticus''), also known as the Mesopotamian softshell turtle, is a species of softshell turtle in the family Trionychidae. It is found throughout much of the Euphrates–Tigris river basin in Iraq, Syria, Turkey and Khūzestān Province of Iran. Historically it has also been reported from Israel, but this likely involves confusion with the very similar '' Trionyx triunguis'' (the two typically require in-hand examination to be separated). The Euphrates softshell turtle is an endangered species that primarily is threatened by habitat loss and alteration, especially the building of dams, but to lesser extent also pollution and killing by fishermen. History The Euphrates softshell turtle became known to western science when the French naturalist Guillaume-Antoine Olivier shot a specimen while crossing the Euphrates near Anah in June 1797. The local residents told him that the meat of this animal was not good to eat, but its fat was ...
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Yangtze Giant Softshell Turtle
The Yangtze giant softshell turtle (''Rafetus swinhoei''), also commonly known as the Red River giant softshell turtle, the Shanghai softshell turtle, the speckled softshell turtle, and Swinhoe's softshell turtle, is a critically endangered species of turtle in the family Trionychidae. It is regarded as one of the largest living freshwater turtle species. Native to eastern and southern China and northern Vietnam, the species has a known population of only two or three individuals, and the absence of fertile females has rendered it functionally extinct. Description The Yangtze giant softshell turtle is recognized for its deep head, pig-like snout, and dorsally positioned eyes. As a critically endangered species, it is widely regarded as the largest freshwater turtle in the world. Although comparisons are limited due to the small number of known specimens, its average and maximum sizes surpass those of other large freshwater turtles, such as the alligator snapping turtle and ...
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Congener (biology)
Biological specificity is the tendency of a characteristic such as a behavior or a biochemical variation to occur in a particular species. Biochemist Linus Pauling stated that "Biological specificity is the set of characteristics of living organisms or constituents of living organisms of being special or doing something special. Each animal or plant species is special. It differs in some way from all other species...biological specificity is the major problem about understanding life." Biological specificity within ''Homo sapiens'' ''Homo sapiens'' has many characteristics that show the biological specificity in the form of behavior and morphology (biology), morphological traits. Morphologically, humans have an enlarged cranial capacity and more gracile features in comparison to other Hominini, hominins. The reduction of dentition is a feature that allows for the advantage of adaptability in diet and survival. As a species, humans are culture dependent and much of human survival ...
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Internal Nares
The choanae (: choana), posterior nasal apertures or internal nostrils are two openings found at the back of the nasal passage between the nasal cavity and the pharynx, in humans and other mammals (as well as crocodilians and most skinks). They are considered one of the most important synapomorphies of tetrapodomorphs, that allowed the passage from water to land. In animals with secondary palates, they allow breathing when the mouth is closed. Janvier, Philippe (2004) "Wandering nostrils". ''Nature'', 432 (7013): 23–24. In tetrapods without secondary palates their function relates primarily to olfaction (sense of smell). The choanae are separated in two by the vomer. Boundaries A choana is the opening between the nasal cavity and the nasopharynx. It is therefore not a structure but a space bounded as follows: * anteriorly and inferiorly by the horizontal plate of palatine bone, * superiorly and posteriorly by the sphenoid bone * laterally by the medial pterygoid plates. ...
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Syntype
In biological nomenclature, a syntype is any one of two or more biological types that is listed in a description of a taxon where no holotype was designated. Precise definitions of this and related terms for types have been established as part of the International Code of Zoological Nomenclature and the International Code of Nomenclature for algae, fungi, and plants. In zoology In zoological nomenclature, a syntype is defined as "Each specimen of a type series (q.v.) from which neither a holotype nor a lectotype has been designated rts. 72.1.2, 73.2, 74 The syntypes collectively constitute the name-bearing type." (Glossary of the zoological Code). Historically, it was common to describe a new species or subspecies from several syntypes without designating a holotype, but this practice is generally frowned upon by modern taxonomists, and most are gradually being replaced by lectotypes. Those that still exist are still considered name-bearing types. A lectotype may be designated ...
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Priority (biology)
Priority is a principle in biological taxonomy by which a valid scientific name is established based on the oldest available name. It is a decisive rule in botanical and zoological nomenclature to recognise the first binomial name (also called ''binominal name'' in zoology) given to an organism as the correct and acceptable name. The purpose is to select one scientific name as a stable one out of two or more alternate names that often exist for a single species. The ''International Code of Nomenclature for algae, fungi, and plants'' (ICN) defines it as: "A right to precedence established by the date of valid publication of a legitimate name or of an earlier homonym, or by the date of designation of a type." Basically, it is a scientific procedure to eliminate duplicate or multiple names for a species, for which Lucien Marcus Underwood called it "the principle of outlaw in nomenclature". History The principle of priority has not always been in place. When Carl Linnaeus laid t ...
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