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Qianshanosuchus
''Qianshanosuchus'' is a genus of basal crocodyloid from the Paleocene of the Qianshan Basin, China. The fossil material, which includes an incomplete skull and parts of the lower jaw, show various features usually associated with juvenile crocodiles alongside various unique traits that were used to erect a new genus. It is the first and only basal crocodyloid currently known from the Paleocene of China, which had previously only yielded alligatoroids and planocraniids. Its presence in this part of the world and its basal position to species of the genus ''Asiatosuchus'' supports the idea that crocodyloids dispersed from Asia into Europe. ''Qianshanosuchus'' only includes a single species, ''Qianshanosuchus youngi''. History and naming ''Qianshanosuchus'' is known from only a single specimen, IBCAS QS15, an incomplete skull and an associated part of the mandible. Based on the small size of the holotype and several anatomical traits, it is thought to preserve a juvenile specime ...
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Crocodyloidea
Crocodyloidea is one of three superfamilies of crocodilians, the other two being Alligatoroidea and Gavialoidea, and it includes the crocodiles. Crocodyloidea may also include the extinct Mekosuchinae, native to Australasia from the Eocene to the Holocene, although this is disputed. Classification Cladistically, it is defined as ''Crocodylus niloticus'' (the Nile crocodile) and all crocodylians more closely related to ''C. niloticus'' than to either ''Alligator mississippiensis'' (the American alligator) or ''Gavialis gangeticus'' (the gharial). This is a stem-based definition for crocodiles, and is more inclusive than the crown group Crocodylidae. As a crown group, Crocodylidae only includes the last common ancestor of all extant (living) crocodiles and their descendants (living or extinct), whereas Crocodyloidea, as a stem group, also includes more basal extinct crocodile ancestors that are more closely related to living crocodiles than to alligators or gavialids. Wh ...
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Crocodyloid
Crocodyloidea is one of three superfamilies of crocodilians, the other two being Alligatoroidea and Gavialoidea, and it includes the crocodiles. Crocodyloidea may also include the extinct Mekosuchinae, native to Australasia from the Eocene to the Holocene, although this is disputed. Classification Cladistically, it is defined as ''Crocodylus niloticus'' (the Nile crocodile) and all crocodylians more closely related to ''C. niloticus'' than to either ''Alligator mississippiensis'' (the American alligator) or ''Gavialis gangeticus'' (the gharial). This is a stem-based definition for crocodiles, and is more inclusive than the crown group Crocodylidae. As a crown group, Crocodylidae only includes the last common ancestor of all extant (living) crocodiles and their descendants (living or extinct), whereas Crocodyloidea, as a stem group, also includes more basal extinct crocodile ancestors that are more closely related to living crocodiles than to alligators or gavialids. When con ...
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Gavialoidea
Gavialoidea is one of three superfamilies of crocodylians, the other two being Alligatoroidea and Crocodyloidea. Although many extinct species are known, only the gharial ''Gavialis gangeticus'' and the false gharial ''Tomistoma schlegelii'' are alive today, with '' Hanyusuchus'' having become extinct in the last few centuries. Extinct South American gavialoids likely dispersed in the mid Tertiary from Africa and Asia. Fossil remains of the Puerto Rican gavialoid '' Aktiogavialis puertorisensis'' were discovered in a cave located in San Sebastián, Puerto Rico and dated to the Oligocene. This individual is thought to have crossed the Atlantic coming from Africa, indicating that this species was able to withstand saltwater. Classification Gavialoidea is cladistically defined as ''Gavialis gangeticus'' (the gharial) and all crocodylians closer to it than to ''Alligator mississippiensis'' (the American alligator) or ''Crocodylus niloticus'' (the Nile crocodile). This is a stem-ba ...
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Crocodylinae
Crocodylinae is a subfamily of true crocodiles within the family Crocodylidae, and is the sister taxon to Osteolaeminae ( dwarf crocodiles and slender-snouted crocodiles). Taxonomy Crocodylinae was cladistically defined by Christopher Brochu in 1999 as ''Crocodylus niloticus'' (the Nile crocodile) and all crocodylians more closely related to it than to ''Osteolaemus tetraspis'' (the Dwarf crocodile). This is a stem-based definition, and is the sister taxon to Osteolaeminae. Crocodylinae contains the extant genus '' Crocodylus''. It is disputed as to whether is also includes '' Mecistops'' (slender-snouted crocodiles), or the extinct genus Voay. Phylogeny Some morphological studies have recovered '' Mecistops'' as a basal member of Crocodylinae, more closely related to '' Crocodylus'' than to '' Osteolaemus'' and the other members of Osteolaeminae, as shown in the cladogram below. The below cladogram is based on a 2021 study using paleogenomics that extracted DNA from th ...
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Osteolaeminae
Osteolaeminae is a subfamily (biology), subfamily of true crocodiles within the family (biology), family Crocodylidae containing the Osteolaemus, dwarf crocodiles and Mecistops, slender-snouted crocodiles, and is the sister taxon to Crocodylinae. Taxonomy Osteolaeminae was named by Christopher Brochu in 2003 as a subfamily of Crocodylidae separate from Crocodylinae, and is cladistics, cladistically defined as ''Osteolaemus tetraspis'' (the Dwarf crocodile) and all crocodylians more closely related to it than to ''Crocodylus niloticus'' (the Nile crocodile). This is a stem-based taxon, stem-based definition, and is the sister taxon to Crocodylinae. Osteolaeminae contains the two extant taxon, extant genus, genera ''Osteolaemus'' and ''Mecistops'', along with several extinct genera, although the number of extant species within Osteolaeminae is currently in question. Phylogeny The cladogram below is based on two studies that combined morphology (biology), morphological, molecular (D ...
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Mekosuchinae
Mekosuchinae is an extinct clade of crocodilians from the Cenozoic of Australasia. They first appear in the fossil record in the Eocene in Australia, and survived until the arrival of humans: in the Pleistocene in Australia and within the Holocene in the Pacific islands of Fiji, New Caledonia and Vanuatu. Mekosuchine crocodiles are a diverse group. One of the last species, '' Mekosuchus inexpectatus'' from Holocene New Caledonia, may have been arboreal. The early Miocene species '' Harpacochampsa camfieldensis'' may have resembled a false gharial. Another mekosuchine fossil, currently undescribed, has been found in Miocene deposits from New Zealand. One genus, '' Mekosuchus'', managed to spread to the islands of the Pacific; it is believed to have island-hopped across the Coral Sea, moving first to a now submerged island known as Greater Chesterfield Island, then New Caledonia and onwards. In the Pleistocene, ''Quinkana'' was one of the top terrestrial predators of the Austra ...
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Asiatosuchus Grangeri
''Asiatosuchus'' is an extinct genus of crocodyloid crocodilian Crocodilia (or Crocodylia, both ) is an order of mostly large, predatory, semiaquatic reptiles, known as crocodilians. They first appeared 95 million years ago in the Late Cretaceous period ( Cenomanian stage) and are the closest living ...s that lived in Eurasia during the Paleogene. Many Paleogene crocodilians from Europe and Asia have been attributed to ''Asiatosuchus'' since the genus was named in 1940. These species have a generalized crocodilian morphology (biology), morphology typified by flat, triangular skulls. The feature that traditionally united these species under the genus ''Asiatosuchus'' is a broad connection or Symphysis menti, symphysis between the two halves of the lower jaw. Recent studies of the evolutionary relationships of early crocodilians along with closer examinations of the morphology of fossil specimens suggest that only the first named species of ''Asiatosuchus'', ''A. grange ...
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Prodiplocynodon Langi
''Prodiplocynodon'' is an extinct genus of basal crocodyloid crocodylian. It is one of the only crocodyloids known from the Cretaceous and existed during the Maastrichtian stage. The only species of ''Prodiplocynodon'' is the type species ''P. langi'' from the Lance Formation of Wyoming, known only from a single holotype skull lacking the lower jaw. The skull was collected by the American Museum Expedition of 1892 from exposures near the Cheyenne River in Niobrara County. It was described by Charles C. Mook of the American Museum of Natural History in 1941. The generic name means "before ''Diplocynodon''" because Mook saw close similarities between the holotype skull and that of the alligatoroid '' Diplocynodon'' from the Eocene of Europe. Description Most of the cranial sutures that outline individual bones of the skull are not visible in the holotype, and are often obscured by cracks. However, the overall shape of ''Prodiplocynodon'' is similar to that of basal all ...
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Albertosuchus
''Albertosuchus'' is an extinct genus of crocodyloid crocodylian from the Late Cretaceous of Canada. The type species ''Albertosuchus knudsenii'' was named in 2015 from the Scollard Formation in Alberta. ''Albertosuchus'' is the northernmost-known Late Cretaceous crocodylian in North America. ''Albertosuchus'' lacks the notch in the upper jaw between the maxilla and premaxilla bones that is characteristic of most crocodyloids, and it also has a very short mandibular symphysis (the connection between the two halves of the lower jaw). Phylogenetic analysis indicates that it is one of the most basal members of Crocodyloidea Crocodyloidea is one of three superfamilies of crocodilians, the other two being Alligatoroidea and Gavialoidea, and it includes the crocodiles. Crocodyloidea may also include the extinct Mekosuchinae, native to Australasia from the Eocene to ... and a close relative of '' Arenysuchus'' from the Late Cretaceous of Spain, although the incomplete nature of kn ...
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Borealosuchus
''Borealosuchus'' (meaning "boreal crocodile") is an extinct genus of crocodyliforms that lived from the Late Cretaceous to the Eocene in North America. It was named by Chris Brochu in 1997 for several species that had been assigned to ''Leidyosuchus''. The species assigned to it are: ''B. sternbergii'', the type species, from the Maastrichtian (Late Cretaceous) of Colorado, Montana, North Dakota, South Dakota, and Wyoming; ''B. acutidentatus'', from the Paleocene of Saskatchewan; ''B. formidabilis'', from the Paleocene of North Dakota; ''B. griffithi'', from the Paleocene of Alberta; and ''B. wilsoni'', from the Eocene of Wyoming. ''B. formidabilis'' is particularly well-known, represented by the remains of many individuals from the Wannagan Creek site in North Dakota. ''Borealosuchus'' was a mid-sized crocodyliform; ''B. acutidentatus'' reached up to in length with a skull. Taxonomy Six species of ''Borealosuchus'' are currently recognized. In order of their naming, the ...
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Crocodilia
Crocodilia (or Crocodylia, both ) is an order of mostly large, predatory, semiaquatic reptiles, known as crocodilians. They first appeared 95 million years ago in the Late Cretaceous period ( Cenomanian stage) and are the closest living relatives of birds, as the two groups are the only known survivors of the Archosauria. Members of the order's total group, the clade Pseudosuchia, appeared about 250 million years ago in the Early Triassic period, and diversified during the Mesozoic era. The order Crocodilia includes the true crocodiles (family Crocodylidae), the alligators and caimans (family Alligatoridae), and the gharial and false gharial (family Gavialidae). Although the term 'crocodiles' is sometimes used to refer to all of these, crocodilians is a less ambiguous vernacular term for members of this group. Large, solidly built, lizard-like reptiles, crocodilians have long flattened snouts, laterally compressed tails, and eyes, ears, and nostrils ...
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Longirostres
Longirostres is a clade of crocodilians that includes the crocodiles and the gavialids, to the exclusion of the alligatoroids. Named in 2003 by Harshman ''et al.'', Longirostres is a crown group defined phylogenetically as including the last common ancestor of ''Crocodylus niloticus'' and '' Gavialis gangeticus'' and all of its descendants. Traditionally, crocodiles and alligators were considered more closely related and grouped together in the clade Brevirostres, to the exclusion of the gharial The gharial (''Gavialis gangeticus''), also known as gavial or fish-eating crocodile, is a crocodilian in the family Gavialidae and among the longest of all living crocodilians. Mature females are long, and males . Adult males have a distinct ...s. This classification was based on morphological studies primarily focused on analyzing skeletal traits of living and extinct fossil species. However, recent molecular studies using DNA sequencing have rejected Brevirostres upon fin ...
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