Pucciniomycetes
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Pucciniomycetes
The Pucciniomycetes (formerly known as the Urediniomycetes) are a class of fungi in the Pucciniomycotina subdivision of the Basidiomycota. The class contains 5 orders, 21 families, 190 genera, and 8016 species. It includes several important plant pathogens causing forms of fungal rust. Characteristics Pucciniomycetes develop no basidiocarp, karyogamy occurs in a thick-walled resting spore ( teliospore), and meiosis occurs upon germination of teliospore. They have simple septal pores without membrane caps and disc-like spindle pole bodies. Except for a few species, the basidia, when present, are transversally septate. Mannose is the major cell wall carbohydrate, glucose, fucose and rhamnose Rhamnose (Rha, Rham) is a naturally occurring deoxy sugar. It can be classified as either a methyl-pentose or a 6-deoxy-hexose. Rhamnose predominantly occurs in nature in its L-form as L-rhamnose (6-deoxy-L-mannose). This is unusual, since most o ... are the less prevalent ne ...
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Helicobasidiales
The Helicobasidiales are an order of rust fungi in the class Pucciniomycetes. It contains the single family Helicobasidiaceae, which itself comprises three genera: ''Helicobasidium'', '' Stypinella'', and '' Tuberculina''. Helicobasidiales was circumscribed In geometry, the circumscribed circle or circumcircle of a polygon is a circle that passes through all the vertices of the polygon. The center of this circle is called the circumcenter and its radius is called the circumradius. Not every po ... in 2006. References Basidiomycota orders Monotypic fungus taxa Taxa named by Franz Oberwinkler Taxa described in 2006 {{Basidiomycota-stub ...
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Platygloeales
The Platygloeales are an order of rust fungi in the class Pucciniomycetes. It contains two families, the '' Eocronartiaceae'' and also the '' Platygloeaceae''. Order Platygloeales are monophyletic group (they have a common ancestor). The results of a molecular analyses investigation in 2007, revealed that the former placement of '' Auriculoscypha'' genus in Platygloeales as in the Dictionary of the Fungi (as per Hawksworth et al. 1995; Kirk et al. 2001) can no longer be justified. Description Order Platygloeales are parasitic on mosses and other plants. They are saprobes on wood or they parasitize other fungi, ferns, mosses or other vascular plants. They have pycnium (fruiting body of rusts) which forms masses of hyphae (long, branching and filamentous structure) inside mosses; example genera include ''Platygloea'' and ''Eocronartium''. As part of Order Pucciniales they typically have 5 spore stages and 2 alternate hosts. They generally have four-celled ''auriculariod'' basidia ...
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Pucciniomycotina
Pucciniomycotina is a subdivision of fungus within the division Basidiomycota. The subdivision contains 9 classes, 20 orders, and 37 families. Over 8400 species of Pucciniomycotina have been described - more than 8% of all described fungi. The subdivision is considered a sister group to Ustilaginomycotina and Agaricomycotina, which may share the basal lineage of Basidiomycota, although this is uncertain due to low support for placement between the three groups. The group was known as Urediniomycetes until 2006, when it was elevated from a class to a subdivision and named after the largest order in the group, Pucciniales. Ecology Pucciniomycotina have a diverse range of ecologies as insect parasites, mycoparasites, and orchid mycorrhiza; some have been detected in soil and water or asymptotic members living on leaves. Most are plant pathogens. Many Pucciniomycotina are rust fungi and are placed in the order Puccinales that contains roughly 7800 species (c. 90% of the group). ...
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Basidiomycota
Basidiomycota () is one of two large divisions that, together with the Ascomycota, constitute the subkingdom Dikarya (often referred to as the "higher fungi") within the kingdom Fungi. Members are known as basidiomycetes. More specifically, Basidiomycota includes these groups: mushrooms, puffballs, stinkhorns, bracket fungi, other polypores, jelly fungi, boletes, chanterelles, earth stars, smuts, bunts, rusts, mirror yeasts, and ''Cryptococcus'', the human pathogenic yeast. Basidiomycota are filamentous fungi composed of hyphae (except for basidiomycota-yeast) and reproduce sexually via the formation of specialized club-shaped end cells called basidia that normally bear external meiospores (usually four). These specialized spores are called basidiospores. However, some Basidiomycota are obligate asexual reproducers. Basidiomycota that reproduce asexually (discussed below) can typically be recognized as members of this division by gross similarity to others, by the form ...
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Fungi
A fungus (plural, : fungi or funguses) is any member of the group of Eukaryote, eukaryotic organisms that includes microorganisms such as yeasts and Mold (fungus), molds, as well as the more familiar mushrooms. These organisms are classified as a Kingdom (biology), kingdom, separately from the other eukaryotic kingdoms, which by one traditional classification include Plantae, Animalia, Protozoa, and Chromista. A characteristic that places fungi in a different kingdom from plants, bacteria, and some protists is chitin in their cell walls. Fungi, like animals, are heterotrophs; they acquire their food by absorbing dissolved molecules, typically by secreting digestive enzymes into their environment. Fungi do not photosynthesize. Growth is their means of motility, mobility, except for spores (a few of which are flagellated), which may travel through the air or water. Fungi are the principal decomposers in ecological systems. These and other differences place fungi in a single gro ...
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Meiosis
Meiosis (; , since it is a reductional division) is a special type of cell division of germ cells in sexually-reproducing organisms that produces the gametes, such as sperm or egg cells. It involves two rounds of division that ultimately result in four cells with only one copy of each chromosome ( haploid). Additionally, prior to the division, genetic material from the paternal and maternal copies of each chromosome is crossed over, creating new combinations of code on each chromosome. Later on, during fertilisation, the haploid cells produced by meiosis from a male and female will fuse to create a cell with two copies of each chromosome again, the zygote. Errors in meiosis resulting in aneuploidy (an abnormal number of chromosomes) are the leading known cause of miscarriage and the most frequent genetic cause of developmental disabilities. In meiosis, DNA replication is followed by two rounds of cell division to produce four daughter cells, each with half the number ...
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Septal Pore
In algal anatomy, a pit connection is a hole in the septum between two algal cells, and is found only in the red algae − specifically, all orders except the Porphyridiales and haploid Bangiales. They are often stoppered with proteinaceous "pit plugs". By contrast, many fungi (only ascomycetes and basidiomycetes, as most other groups lack septa) contain septal pores − an unrelated phenomenon. Characteristics A sieve-like membrane may cover the pit in living algae, but in the majority of algae a plug forms, they likely limit the transfer of metabolites between neighbouring cells. Formation Primary pit connections are formed between cells in the same filament, derived from the same parent cell by its division. Such connections are always single, and usually circular; this is a result of their method of formation. The septum is formed as the walls of a filament grow inwards, dividing the cell; this results in a hole in the middle of the tube where the walls don't quite merge. ...
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Mitotic Spindle
In cell biology, the spindle apparatus refers to the cytoskeletal structure of eukaryotic cells that forms during cell division to separate sister chromatids between daughter cells. It is referred to as the mitotic spindle during mitosis, a process that produces genetically identical daughter cells, or the meiotic spindle during meiosis, a process that produces gametes with half the number of chromosomes of the parent cell. Besides chromosomes, the spindle apparatus is composed of hundreds of proteins. Microtubules comprise the most abundant components of the machinery. Spindle structure Attachment of microtubules to chromosomes is mediated by kinetochores, which actively monitor spindle formation and prevent premature anaphase onset. Microtubule polymerization and depolymerization dynamic drive chromosome congression. Depolymerization of microtubules generates tension at kinetochores; bipolar attachment of sister kinetochores to microtubules emanating from opposi ...
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Microtubule Organizing Center
The microtubule-organizing center (MTOC) is a structure found in eukaryotic cells from which microtubules emerge. MTOCs have two main functions: the organization of eukaryotic flagella and cilia and the organization of the mitotic and meiotic spindle apparatus, which separate the chromosomes during cell division. The MTOC is a major site of microtubule nucleation and can be visualized in cells by immunohistochemical detection of γ-tubulin. The morphological characteristics of MTOCs vary between the different phyla and kingdoms. In animals, the two most important types of MTOCs are 1) the basal bodies associated with cilia and flagella and 2) the centrosome associated with spindle formation. Organization Microtubule-organizing centers function as the site where microtubule formation begins, as well as a location where free-ends of microtubules attract to. Within the cells, microtubule-organizing centers can take on many different forms. An array of microtubules can arrange t ...
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Basidium
A basidium () is a microscopic sporangium (a spore-producing structure) found on the hymenophore of fruiting bodies of basidiomycete fungi which are also called tertiary mycelium, developed from secondary mycelium. Tertiary mycelium is highly-coiled secondary myceliuma dikaryon. The presence of basidia is one of the main characteristic features of the Basidiomycota. A basidium usually bears four sexual spores called basidiospores; occasionally the number may be two or even eight. In a typical basidium, each basidiospore is borne at the tip of a narrow prong or horn called a sterigma (), and is forcibly discharged upon maturity. The word ''basidium'' literally means "little pedestal", from the way in which the basidium supports the spores. However, some biologists suggest that the structure more closely resembles a club. An immature basidium is known as a basidiole. Structure Most basidiomycota have single celled basidia (holobasidia), but in some groups basidia can be ...
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Puccinia Sessilis
''Puccinia sessilis'' is a fungal plant pathogen, also known as arum rust or ramsons rust. It commonly infects ''Arum maculatum'' and ''Allium ursinum'' causing yellow to orange circular patches on leaves. On the underside it produces raised orange aecia commonly covered in spores. It is common in Eurasia in the spring. Other species affected by this rust include ''Convallaria majalis'', ''Dactylorhiza fuchsii'', ''Dactylorhiza incarnata'', ''Dactylorhiza majalis'', ''Gymnadenia conopsea'', ''Neottia ovata'', ''Paris quadrifolia'' and ''Phalaris arundinacea'' A specialised form, ''Puccinia sessilis'' f.sp. ''narcissi-orchidacearum'' Boerema & Kesteren (''Aecidium narcissi'') is a cause of rust in daffodils ('' Narcissus'') and various wild Orchidaceae Orchids are plants that belong to the family Orchidaceae (), a diverse and widespread group of flowering plants with blooms that are often colourful and fragrant. Along with the Asteraceae, they are one of the two largest fa ...
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Mannose
Mannose is a sugar monomer of the aldohexose series of carbohydrates. It is a C-2 epimer of glucose. Mannose is important in human metabolism, especially in the glycosylation of certain proteins. Several congenital disorders of glycosylation are associated with mutations in enzymes involved in mannose metabolism. Mannose is not an essential nutrient; it can be produced in the human body from glucose, or converted into glucose. Mannose provides 2–5 kcal/g. It is partially excreted in the urine. Etymology The root of both "mannose" and "mannitol" is manna, which the Bible describes as the food supplied to the Israelites during their journey in the region of Sinai. Several trees and shrubs can produce a substance called manna, such as the "manna tree" ('' Fraxinus ornus'') from whose secretions mannitol was originally isolated. Structure Mannose commonly exists as two different-sized rings, the pyranose (six-membered) form and the furanose (five-membered) form. Eac ...
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