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Pseudocoelomata
The Aschelminthes (Aeschelminthes or Nemathelminthes), closely associated with the Platyhelminthes, are an obsolete phylum of pseudocoelomate and other similar animals that are no longer considered closely related and have been promoted to phyla in their own right. The term Aschelminth is now generally only used as an informal name for any member of the approximately ten different invertebrate phyla formerly included within Aschelminthes. It is a polyphyletic group. Subdivisions Although invertebrate experts do not necessarily agree on these categorizations, groups that are generally incorporated into Aschelminthes include: * Gastrotricha Gnathifera * Acanthocephala * Gnathostomulida * Chaetognatha * Rotifera Nematoidea * Nematoda * Nematomorpha Scalidophora * Kinorhyncha * Loricifera * Priapulida In addition, Cycliophora, Entoprocta and Tardigrada Tardigrades (), known colloquially as water bears or moss piglets, are a phylum of eight-legged Segmentation (biology), ...
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Nematode
The nematodes ( or ; ; ), roundworms or eelworms constitute the phylum Nematoda. Species in the phylum inhabit a broad range of environments. Most species are free-living, feeding on microorganisms, but many are parasitic. Parasitic worms (helminths) are the cause of soil-transmitted helminthiases. They are classified along with arthropods, tardigrades and other moulting animals in the clade Ecdysozoa. Unlike the flatworms, nematodes have a tubular digestive system, with openings at both ends. Like tardigrades, they have a reduced number of Hox genes, but their sister phylum Nematomorpha has kept the ancestral protostome Hox genotype, which shows that the reduction has occurred within the nematode phylum. Nematode species can be difficult to distinguish from one another. Consequently, estimates of the number of nematode species are uncertain. A 2013 survey of animal biodiversity suggested there are over 25,000. Estimates of the total number of extant species are su ...
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Pseudocoelomate
The coelom (or celom) is the main body cavity in many animals and is positioned inside the body to surround and contain the digestive tract and other organs. In some animals, it is lined with mesothelium. In other animals, such as molluscs, it remains undifferentiated. In the past, and for practical purposes, coelom characteristics have been used to classify bilaterian animal phyla into informal groups. Etymology The term ''coelom'' derives from the Ancient Greek word () 'cavity'. Structure Development The coelom is the mesodermally lined cavity between the gut and the outer body wall. During the development of the embryo, coelom formation begins in the gastrulation stage. The developing digestive tube of an embryo forms as a blind pouch called the archenteron. In protostomes, the coelom forms by a process known as schizocoely. The archenteron initially forms, and the mesoderm splits into two layers: the first attaches to the body wall or ectoderm, forming the parieta ...
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Platyhelminthes
Platyhelminthes (from the Greek πλατύ, ''platy'', meaning "flat" and ἕλμινς (root: ἑλμινθ-), ''helminth-'', meaning "worm") is a phylum of relatively simple bilaterian, unsegmented, soft-bodied invertebrates commonly called flatworms or flat worms. Being acoelomates (having no body cavity), and having no specialised circulatory and respiratory organs, they are restricted to having flattened shapes that allow oxygen and nutrients to pass through their bodies by diffusion. The digestive cavity has only one opening for both ingestion (intake of nutrients) and egestion (removal of undigested wastes); as a result, the food can not be processed continuously. In traditional medicinal texts, Platyhelminthes are divided into Turbellaria, which are mostly non- parasitic animals such as planarians, and three entirely parasitic groups: Cestoda, Trematoda and Monogenea; however, since the turbellarians have since been proven not to be monophyletic, this classification i ...
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Entoprocta
Entoprocta (), or Kamptozoa , is a phylum (biology), phylum of mostly Sessility (zoology), sessile aquatic animals, ranging from long. Mature individuals are goblet-shaped, on relatively long stalks. They have a "crown" of solid tentacles whose cilia generate water currents that filter feeder, draw food particles towards the mouth, and both the mouth and anus lie inside the "crown". The superficially similar Bryozoa (Ectoprocta) have the anus outside a "crown" of hollow tentacles. Most family (biology), families of entoprocts are colonial, and all but 2 of the 150 species are marine. A few solitary species can move slowly. Some species eject unfertilized ovum, ova into the water, while others keep their ova in brood chambers until they hatch, and some of these species use placenta-like organs to nourish the developing eggs. After hatching, the larvae swim for a short time and then settle on a surface. There they Metamorphosis, metamorphose, and the larval gut rotates by up ...
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Cycliophora
''Symbion'' is a genus of commensal aquatic animals, less than 0.5 mm wide, found living attached to the mouthparts of cold-water lobsters. They have sac-like bodies, and three distinctly different forms in different parts of their two-stage life-cycle. They appear so different from other animals that they were assigned their own, new phylum Cycliophora shortly after they were discovered in 1995. This was the first new phylum of multicelled organism to be discovered since the Loricifera in 1983. Taxonomy ''Symbion pandora'' was discovered in 1995 by Reinhardt Kristensen and Peter Funch on the mouthparts of the Norway lobster (''Nephrops norvegicus''). Other, related, species have since been discovered on: * the American lobster (''Homarus americanus'', host to '' Symbion americanus'') * the European lobster (''Homarus gammarus'', host to an as yet unnamed species of ''Symbion'') The genus is so named because of its commensal relationship with the lobster (a form ...
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Priapulida
Priapulida (priapulid worms, from Gr. πριάπος, ''priāpos'' ' Priapus' + Lat. ''-ul-'', diminutive), sometimes referred to as penis worms, is a phylum of unsegmented marine worms. The name of the phylum relates to the Greek god of fertility, because their general shape and their extensible spiny introvert (eversible) proboscis may resemble the shape of a human penis. They live in the mud, except for a few tropical meiobenthic species which live in medium- to coarse-grained sands, and are found in comparatively shallow waters to deep waters and no warmer than 12–13°C. Some species show a remarkable tolerance for hydrogen sulfide, anoxia and low salinity. ''Halicryptus spinulosus'' appears to prefer brackish shallow waters. They can be quite abundant in some areas. In an Alaskan bay as many as 85 adult individuals of '' Priapulus caudatus'' per square meter has been recorded, while the density of its larvae can be as high as 58,000 per square meter (5,390 per square foot). ...
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Loricifera
Loricifera (from Latin, ''wikt:lorica, lorica'', corselet (armour) + ''ferre'', to bear) is a phylum of very small to microscopic marine cycloneuralian sediment-dwelling animals with 43 described species and approximately 100 more that have been collected and not yet described. Their sizes range from 100 μm to . They are characterised by a protective outer case called a Lorica (biology), lorica and their habitat is in the spaces between marine gravel to which they attach themselves. The phylum was discovered in 1983 by Reinhardt Kristensen, R.M. Kristensen, near Roscoff, France. They are among the most recently discovered groups of animals. They attach themselves quite firmly to the substrate, and hence remained undiscovered for so long. The first specimen was collected in the 1970s, and described in 1983. They are found at all depths, in different sediment types, and in all latitudes. Morphology The animals have a head, mouth, and digestive system, as well as the lo ...
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Kinorhyncha
Kinorhyncha (, ' "snout") is a phylum of small marine invertebrates that are widespread in mud or sand at all depths as part of the meiobenthos. They are commonly called mud dragons. Modern species are or less, but Cambrian forms could reach . Anatomy Kinorhynchs are limbless animals, with a body consisting of a head, neck, and a trunk of eleven segments. They are the only members of Ecdysozoa, except from the panarthropoda, with a segmented body. Juveniles have eight or nine segments, depending on genus, with the last two or three being added later during growth. A Cambrian species, Eokinorhynchus rarus, had about twice as many segments as present forms. Like other ecdysozoans they do not have external cilia, but instead have a number of spines along the body, plus up to seven circles of spines around the head. These spines are used for locomotion, withdrawing the head and pushing forward, then gripping the substrate with the spines while drawing up the body. The body ...
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Scalidophora
Scalidophora is a group of marine pseudocoelomate ecdysozoans that was proposed on morphological grounds to unite three phyla: the Kinorhyncha, the Priapulida and the Loricifera. The three phyla have four characters in common — chitinous cuticle that is moulted, rings of scalids on the introvert, flosculi, and two rings of introvert retracts.Heiner, I., Kristensen, R.H. 2005. Two new species of the genus ''Pliciloricus'' (Loricifera, Pliciloricidae) from the Faroe Bank, North Atlantic. Zoologischer Anzeiger. 243: 121–138. The introvert and abdomen are separated by a distinct neck region in all groups, but in adult macroscopic priapulids it becomes rudimentary in ''Priapulus'' and is completely absent in ''Halicryptus''. However, the monophyly of the Scalidophora was not supported by two molecular studies, where the position of the Loricifera was uncertain or as sister to the Panarthropoda. Both studies supported a reduced Scalidophora comprising the Kinorhyncha and Priapulid ...
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Nematomorpha
Nematomorpha (sometimes called Gordiacea, and commonly known as horsehair worms, hairsnakes, or Gordian worms) are a phylum of parasitoid animals superficially similar to nematode worms in morphology, hence the name. Most species range in size from , reaching in extreme cases, and in diameter. Horsehair worms can be discovered in damp areas, such as watering troughs, swimming pools, streams, puddles, and cisterns. The adult worms are free-living, but the larvae are parasitic on arthropods, such as beetles, cockroaches, mantises, orthopterans, and crustaceans. About 351 freshwater species are known and a conservative estimate suggests that there may be about 2000 freshwater species worldwide. The name "Gordian" stems from the legendary Gordian knot. This relates to the fact that nematomorphs often coil themselves in tight balls that resemble knots. Description and biology Nematomorphs possess an external cuticle without cilia. Internally, they have only longitudinal muscle and ...
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Nematoidea
Nematoida is a clade of pseudocoelomate free living or parasitic animals. It consists of phyla Nematoda and Nematomorpha. The two groups share a number of features in common; the presence of a cloaca in both sexes, aflagellate sperm, and a cuticle made of collagen. Its position within Ecdysozoa is uncontroversial, but the identity of their closest relatives has been debated. Under the Cycloneuralia hypothesis, the nematoids are considered to be closest to Scalidophora Scalidophora is a group of marine pseudocoelomate ecdysozoans that was proposed on morphological grounds to unite three phyla: the Kinorhyncha, the Priapulida and the Loricifera. The three phyla have four characters in common — chitinous cuti ..., named for the ring-shaped brains found in these animals. However, this group has seen less support in phylogenetic analyses, with the Cryptovermes hypothesis being more consistently supported, which groups the nematoids with the panarthropods, although the issue ...
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Rotifer
The rotifers (, from Latin 'wheel' and 'bearing'), sometimes called wheel animals or wheel animalcules, make up a phylum (Rotifera ) of microscopic and near-microscopic Coelom#Pseudocoelomates, pseudocoelomate animals. They were first described by John Harris (writer), Rev. John Harris in 1696, and other forms were described by Antonie van Leeuwenhoek in 1703. Most rotifers are around long (although their size can range from to over ), and are common in freshwater environments throughout the world with a few Seawater, saltwater species. Some rotifers are free swimming and truly planktonic, others move by inchworming along a substrate, and some are Sessility (zoology), sessile, living inside tubes or gelatinous holdfast (biology), holdfasts that are attached to a substrate. About 25 species are colonial (e.g., ''Sinantherina semibullata''), either sessile or planktonic. Rotifers are an important part of the freshwater zooplankton, being a major foodsource and with many specie ...
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