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Protungulatum Donnae
''Protungulatum'' ('first ungulate') is an extinct genus of eutherian mammals within extinct family Protungulatidae, and is possibly one of the earliest known placental mammals in the fossil record, that lived in North America from the Late Cretaceous to early Paleocene.Maeva Orliac (2016."The inner ear of ''Protungulatum'' (Pan-Euungulata, Mammalia)"Journal of Mammalian Evolution 23(4) Fossils of this genus were first found in the Bug Creek Anthills in northeastern Montana. The Bug Creek Anthills were initially believed to be Late Cretaceous (latest Maastrichtian) because of the presence of the remains of non-avian dinosaurs and common Cretaceous mammals, but these were later shown to have been reworked from Late Cretaceous strata, and consequently the Bug Creek Anthills are currently believed to be Early Paleocene (Puercan) in age. Remains from the Ravenscrag Formation of Saskatchewan, Canada have been assigned to ''Protungulatum donnae''. These remains may also be Cretaceo ...
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Maastrichtian
The Maastrichtian ( ) is, in the International Commission on Stratigraphy (ICS) geologic timescale, the latest age (geology), age (uppermost stage (stratigraphy), stage) of the Late Cretaceous epoch (geology), Epoch or Upper Cretaceous series (stratigraphy), Series, the Cretaceous geologic period, Period or system (stratigraphy), System, and of the Mesozoic geologic era, Era or Erathem. It spanned the interval from . The Maastrichtian was preceded by the Campanian and succeeded by the Danian (part of the Paleogene and Paleocene). It is named after the city of Maastricht, the capital and largest city of the Limburg (Netherlands), Limburg province in the Netherlands. The Cretaceous–Paleogene extinction event (formerly known as the Cretaceous–Tertiary period, Tertiary extinction event) occurred at the end of this age. In this extinction event, mass extinction, many commonly recognized groups such as non-avian dinosaurs, plesiosaurs and mosasaurs, as well as many other lesser-kn ...
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Strata
In geology and related fields, a stratum (: strata) is a layer of Rock (geology), rock or sediment characterized by certain Lithology, lithologic properties or attributes that distinguish it from adjacent layers from which it is separated by visible surfaces known as either ''Bed (geology), bedding surfaces'' or ''bedding planes''.Salvador, A. ed., 1994. ''International stratigraphic guide: a guide to stratigraphic classification, terminology, and procedure. 2nd ed.'' Boulder, Colorado, The Geological Society of America, Inc., 215 pp. . Prior to the publication of the International Stratigraphic Guide, older publications have defined a stratum as being either equivalent to a single Bed (geology), bed or composed of a number of beds; as a layer greater than 1 cm in thickness and constituting a part of a bed; or a general term that includes both ''bed'' and ''Lamination (geology), lamina''.Neuendorf, K.K.E., Mehl, Jr., J.P., and Jackson, J.A. , eds., 2005. ''Glossary of Geolo ...
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Science (journal)
''Science'' is the peer review, peer-reviewed academic journal of the American Association for the Advancement of Science (AAAS) and one of the world's top academic journals. It was first published in 1880, is currently circulated weekly and has a subscriber base of around 130,000. Because institutional subscriptions and online access serve a larger audience, its estimated readership is over 400,000 people. ''Science'' is based in Washington, D.C., United States, with a second office in Cambridge, UK. Contents The major focus of the journal is publishing important original scientific research and research reviews, but ''Science'' also publishes science-related news, opinions on science policy and other matters of interest to scientists and others who are concerned with the wide implications of science and technology. Unlike most scientific journals, which focus on a specific field, ''Science'' and its rival ''Nature (journal), Nature'' cover the full range of List of academ ...
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Pan-Euungulata
Pan-Euungulata (literally: "all true ungulates"; also known as: Euungulata ensu latoref name="Av"/>) is a clade of placental mammals from grandorder Ferungulata, consisting of the taxon Euungulata ensu strictoand all taxa (species) more closely related to it than to any other living species. The specification of the above-mentioned taxa related to Euungulata s.s. varies from author to author and changes over time.Mark E. Samuels, Sophie Regnault, John R. Hutchinson (2017."Evolution of the patellar sesamoid bone in mammals"PeerJ.; 5: e3103Conference abstract (p. 99)
Explanation and conclusions: .
The extinct family Protungulatidae is a typical representative of such taxa.A ...
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Euungulata
Ungulates ( ) are members of the diverse clade Euungulata ("true ungulates"), which primarily consists of large mammals with Hoof, hooves. Once part of the clade "Ungulata" along with the clade Paenungulata, "Ungulata" has since been determined to be a polyphyletic and thereby invalid clade based on molecular data. As a result, true ungulates had since been reclassified to the newer clade Euungulata in 2001 within the clade Laurasiatheria while Paenungulata has been reclassified to a distant clade Afrotheria. Living ungulates are divided into two orders: Perissodactyla including Equidae, equines, rhinoceroses, and tapirs; and Artiodactyla including Bos, cattle, antelope, Sus (genus), pigs, giraffes, camels, Ovis, sheep, deer, and Hippopotamidae, hippopotamuses, among others. Cetaceans such as Whale, whales, Dolphin, dolphins, and Porpoise, porpoises are also classified as artiodactyls, although they do not have hooves. Most terrestrial ungulates use the hoofed tips of their toes ...
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Placentalia
Placental mammals ( infraclass Placentalia ) are one of the three extant subdivisions of the class Mammalia, the other two being Monotremata and Marsupialia. Placentalia contains the vast majority of extant mammals, which are partly distinguished from monotremes and marsupials in that the fetus is carried in the uterus of its mother to a relatively late stage of development. The name is something of a misnomer, considering that marsupials also nourish their fetuses via a placenta, though for a relatively briefer period, giving birth to less-developed young, which are then nurtured for a period inside the mother's pouch. Placentalia represents the only living group within Eutheria, which contains all mammals that are more closely related to placentals than they are to marsupials. Anatomical features Placental mammals are anatomically distinguished from other mammals by: * a sufficiently wide opening at the bottom of the pelvis to allow the birth of a large baby relative to ...
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Polyphyletic
A polyphyletic group is an assemblage that includes organisms with mixed evolutionary origin but does not include their most recent common ancestor. The term is often applied to groups that share similar features known as Homoplasy, homoplasies, which are explained as a result of convergent evolution. The arrangement of the members of a polyphyletic group is called a polyphyly .. [Source for pronunciation.] It is contrasted with monophyly and paraphyly. For example, the biological characteristic of warm-bloodedness evolved separately in the ancestors of mammals and the ancestors of birds; "warm-blooded animals" is therefore a polyphyletic grouping. Other examples of polyphyletic groups are algae, C4 photosynthesis, C4 photosynthetic plants, and Xenarthra#Evolutionary relationships, edentates. Many taxonomists aim to avoid homoplasies in grouping taxa together, with a goal to identify and eliminate groups that are found to be polyphyletic. This is often the stimulus for major re ...
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Condylarthra
Condylarthra is an informal group – previously considered an order – of extinct placental mammals, known primarily from the Paleocene and Eocene epochs. They are considered early, primitive ungulates and is now largely considered to be a wastebasket taxon, having served as a dumping ground for classifying ungulates which had not been clearly established as part of either Perissodactyla or Artiodactyla, being composed thus of several unrelated lineages. Taxonomic history Condylarthra always was a problematic group. When first described by , Phenacodontidae was the type and only family therein. , however, raised Condylarthra to an order and included a wide range of diverse placentals with generalized dentitions and postcranial skeletons. More recent researchers (i.e. post-WW2) have been more restrictive; either including only a limited number of taxa, or proposing that the term should be abandoned altogether. Due to their primitive characteristics condylarths have been consi ...
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Arctocyonidae
Arctocyonidae (from Greek ''arktos'' and ''kyôn'', "bear/dog-like") is an extinct, possibly polyphyletic family of placental mammals which lived from the late Cretaceous to the early Eocene. They were initially regarded as creodonts, though have since been reassigned to an order of their own, the Arctocyonia. Some have suggested that arctocyonids are ancestral to modern-day artiodactyls, or that they form a sister group. However, more recent phylogenetic analyses suggest that arctocyonids may represent an artificial grouping of extinct ungulates, or that they might be an assemblage of unrelated placentals related to pangolins, pantodonts, and periptychids. Members of Arctocyonidae are characterised by long skulls, with large sagittal crests and very large canines. In the case of '' Arctocyon'' proper, the lower canines especially were large enough to require a diastema on the upper jaw to accommodate them. Arctocyonids varied considerably in size and morphology. Smaller gen ...
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Hell Creek Formation
The Hell Creek Formation is an intensively studied division of mostly Upper Cretaceous and some lower Paleocene rocks in North America, named for exposures studied along Hell Creek, near Jordan, Montana. The Formation (stratigraphy), formation stretches over portions of Montana, North Dakota, South Dakota, and Wyoming. In Montana, the Hell Creek Formation overlies the Fox Hills Formation. The site of Pompeys Pillar National Monument is a small isolated section of the Hell Creek Formation. In 1966, the Hell Creek Fossil Area was designated as a National Natural Landmark by the National Park Service. It is a series of fresh and brackish-water clays, mudstones, and sandstones deposited during the Maastrichtian and Danian (respectively, the end of the Cretaceous period and the beginning of the Paleogene) by fluvial activity in fluctuating river channels and deltas and very occasional peaty swamp deposits along the low-lying eastern continental margin fronting the late Cretaceous West ...
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2011 In Paleontology
Protozoa New taxa Plants Ferns and fern allies Gymnosperms Gymnosperm research *An amplified whole plant reconstruction of the Ypresian Princeton chert pine ''Pinus arnoldii'', expanding the diagnosis to include ''P. similkameenensis'' (Miller, 1973 in paleontology, 1973) foliage and wood plus unnamed pollens cones found in attachment to the ''P. arnoldii'' ovulate cones is published by Klymiuk, Stockey, & Rothwell. Angiosperms Nematoda Lobopods Vetulicolians Molluscs Arthropods Fishes Amphibians Newly named lepospondyls Newly named temnospondyls Newly named lissamphibians Basal reptiles Newly named captorhinids Newly named basal diapsids Newly named ichthyosaurs Lepidosauromorphs Newly named saurosphargids Newly named sauropterygians Newly named lizards Newly named snakes Turtles Newly named turtles Archosauromorphs Newly named basal archosauromorphs Archosaurs Synapsids Newly named non-mammalian synapsids Mammals Fo ...
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