Promethearchaeati
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Promethearchaeati
Promethearchaeati ( syn. "Asgard") is a kingdom belonging to the domain Archaea that contain eukaryotic signature proteins. It appears that the eukaryotes, the domain that contains the animals, plants, fungi and protists, emerged within the Promethearchaeati, in a branch containing the "''Candidatus'' Heimdallarchaeota". This supports the two-domain system of classification over the three-domain system. The kingdom contains one phylum Promethearchaeota, and it contains one valid class Promethearchaeia. After including the kingdom category into ICNP, the only validly published name of this group is kingdom Promethearchaeati. All formerly proposed "phyla" would be de-ranked to classes in this framework. Discovery and nomenclature In the summer of 2010, sediments were analysed from a gravity core taken in the rift valley on the Knipovich ridge in the Arctic Ocean, near the Loki's Castle hydrothermal vent site. Specific sediment horizons previously shown to contain high abundance ...
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Archaea
Archaea ( ) is a Domain (biology), domain of organisms. Traditionally, Archaea only included its Prokaryote, prokaryotic members, but this has since been found to be paraphyletic, as eukaryotes are known to have evolved from archaea. Even though the domain Archaea Cladistics, cladistically includes eukaryotes, the term "archaea" (: archaeon , from the Greek "ἀρχαῖον", which means ancient) in English still generally refers specifically to prokaryotic members of Archaea. Archaea were initially Taxonomy (biology), classified as bacteria, receiving the name archaebacteria (, in the Archaebacteria Kingdom (biology), kingdom), but this term has fallen out of use. Archaeal cells have unique properties separating them from Bacteria and Eukaryote, Eukaryota. Archaea are further divided into multiple recognized phylum, phyla. Classification is difficult because most have not been Isolation (microbiology), isolated in a laboratory and have been detected only by their Gene, gene s ...
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Two-domain System
The two-domain system is a biological classification by which all organisms in the tree of life are classified into two domains, Bacteria and Archaea. It emerged from development of knowledge of archaea diversity and challenges the widely accepted three-domain system that classifies life into Bacteria, Archaea, and Eukarya. It was preceded by the eocyte hypothesis of James A. Lake in the 1980s, which was largely superseded by the three-domain system, due to evidence at the time. Better understanding of archaea, especially of their roles in the origin of eukaryotes through symbiogenesis with bacteria, led to the revival of the eocyte hypothesis in the 2000s. The two-domain system became more widely accepted after the discovery of a large kingdom of archaea called Promethearchaeati in 2017, which evidence suggests to be the evolutionary root of eukaryotes, thereby making eukaryotes members of the domain Archaea. While the features of promethearchaea do not completely rule out th ...
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Eukaryote
The eukaryotes ( ) constitute the Domain (biology), domain of Eukaryota or Eukarya, organisms whose Cell (biology), cells have a membrane-bound cell nucleus, nucleus. All animals, plants, Fungus, fungi, seaweeds, and many unicellular organisms are eukaryotes. They constitute a major group of Outline of life forms, life forms alongside the two groups of prokaryotes: the Bacteria and the Archaea. Eukaryotes represent a small minority of the number of organisms, but given their generally much larger size, their collective global biomass is much larger than that of prokaryotes. The eukaryotes emerged within the archaeal Kingdom (biology), kingdom Asgard (Archaea), Promethearchaeati and its sole phylum Promethearchaeota. This implies that there are only Two-domain system, two domains of life, Bacteria and Archaea, with eukaryotes incorporated among the Archaea. Eukaryotes first emerged during the Paleoproterozoic, likely as Flagellated cell, flagellated cells. The leading evolutiona ...
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Eukaryogenesis
Eukaryogenesis, the process which created the eukaryotic cell and lineage, is a milestone in the evolution of life, since eukaryotes include all complex cells and almost all multicellular organisms. The process is widely agreed to have involved symbiogenesis, in which an archaeon and a bacterium came together to create the first eukaryotic common ancestor (FECA). This cell had a new level of complexity and capability, with a nucleus, at least one centriole and cilium, facultatively aerobic mitochondria, sex (meiosis and syngamy), a dormant cyst with a cell wall of chitin and/or cellulose and peroxisomes. It evolved into a population of single-celled organisms that included the last eukaryotic common ancestor (LECA), gaining capabilities along the way, though the sequence of the steps involved has been disputed, and may not have started with symbiogenesis. In turn, the LECA gave rise to the eukaryotes' crown group, containing the ancestors of animals, fungi, plants, and a diverse ...
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Promethearchaeota
Promethearchaeota is the sole phylum of the Kingdom (biology), kingdom Asgard (Archaea), Promethearchaeati, the kingdom from which the eukaryotes emerged. The phylum has one valid Class (biology), class: Promethearchaeia. The phylum includes all members of the group previously named Deep Sea Archaeal Group, also known as Marine Benthic Group B. A Phylogenetics, phylogenetic analysis disclosed a monophyletic grouping of the Promethearchaeota with the eukaryotes. The analysis revealed several genes with cell membrane-related functions. The presence of such genes support the hypothesis of an archaeal host for the emergence of the eukaryotes; the Eocyte hypothesis, eocyte-like scenarios. Promethearchaeota was introduced in 2015 after the identification of a candidate genome in a metagenomics, metagenomic analysis of a mid-oceanic sediment sample. This analysis suggests the existence of a genus of unicellular life dubbed ''Lokiarchaeum''. The sample was taken near a hydrothermal vent ...
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Lokiarchaeota
Promethearchaeota is the sole phylum of the kingdom Promethearchaeati, the kingdom from which the eukaryotes emerged. The phylum has one valid class: Promethearchaeia. The phylum includes all members of the group previously named Deep Sea Archaeal Group, also known as Marine Benthic Group B. A phylogenetic analysis disclosed a monophyletic grouping of the Promethearchaeota with the eukaryotes. The analysis revealed several genes with cell membrane-related functions. The presence of such genes support the hypothesis of an archaeal host for the emergence of the eukaryotes; the eocyte-like scenarios. Promethearchaeota was introduced in 2015 after the identification of a candidate genome in a metagenomic analysis of a mid-oceanic sediment sample. This analysis suggests the existence of a genus of unicellular life dubbed ''Lokiarchaeum''. The sample was taken near a hydrothermal vent at a vent field known as Loki's Castle located at the bend between Mohns/Knipovich ridge in the ...
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Three-domain System
The three-domain system is a taxonomic classification system that groups all cellular life into three domains, namely Archaea, Bacteria and Eukarya, introduced by Carl Woese, Otto Kandler and Mark Wheelis in 1990. The key difference from earlier classifications such as the two-empire system and the five-kingdom classification is the splitting of Archaea (previously named "archaebacteria") from Bacteria as completely different organisms. The three domain hypothesis is considered obsolete by some since it is thought that eukaryotes do not form a separate domain of life; instead, they arose from a fusion between two different species, one from within Archaea and one from within Bacteria. (see Two-domain system) Background Woese argued, on the basis of differences in 16S rRNA genes, that bacteria, archaea, and eukaryotes each arose separately from an ancestor with poorly developed genetic machinery, often called a progenote. To reflect these primary lines of descent, he tr ...
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Phylum
In biology, a phylum (; : phyla) is a level of classification, or taxonomic rank, that is below Kingdom (biology), kingdom and above Class (biology), class. Traditionally, in botany the term division (taxonomy), division has been used instead of phylum, although the International Code of Nomenclature for algae, fungi, and plants accepts the terms as equivalent. Depending on definitions, the animal kingdom Animalia contains about 31 phyla, the plant kingdom Plantae contains about 14 phyla, and the fungus kingdom Fungi contains about eight phyla. Current research in phylogenetics is uncovering the relationships among phyla within larger clades like Ecdysozoa and Embryophyta. General description The term phylum was coined in 1866 by Ernst Haeckel from the Greek (, "race, stock"), related to (, "tribe, clan"). Haeckel noted that species constantly evolved into new species that seemed to retain few consistent features among themselves and therefore few features that distinguishe ...
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Class (biology)
In biological classification, class () is a taxonomic rank, as well as a taxonomic unit, a taxon, in that rank. It is a group of related taxonomic orders. Other well-known ranks in descending order of size are domain, kingdom, phylum, order, family, genus, and species, with class ranking between phylum and order. History The class as a distinct rank of biological classification having its own distinctive name – and not just called a ''top-level genus'' ''(genus summum)'' – was first introduced by French botanist Joseph Pitton de Tournefort in the classification of plants that appeared in his '' Eléments de botanique'' of 1694. Insofar as a general definition of a class is available, it has historically been conceived as embracing taxa that combine a distinct ''grade'' of organization—i.e. a 'level of complexity', measured in terms of how differentiated their organ systems are into distinct regions or sub-organs—with a distinct ''type'' of construction, whic ...
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International Code Of Nomenclature Of Prokaryotes
The International Code of Nomenclature of Prokaryotes (ICNP) or Prokaryotic Code, formerly the International Code of Nomenclature of Bacteria (ICNB) or Bacteriological Code (BC), governs the scientific names for Bacteria and Archaea.P. H. A. Sneath, 2003. A short history of the Bacteriological CodURL It denotes the rules for naming taxa of bacteria, according to their relative rank. As such it is one of the nomenclature codes of biology. Originally the '' International Code of Botanical Nomenclature'' dealt with bacteria, and this kept references to bacteria until these were eliminated at the 1975 International Botanical Congress. An early Code for the nomenclature of bacteria was approved at the 4th International Congress for Microbiology in 1947, but was later discarded. The latest version to be printed in book form is the 1990 Revision, but the book does not represent the current rules. The 2008 and 2022 Revisions have been published in the '' International Journal of Systemati ...
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List Of Prokaryotic Names With Standing In Nomenclature
List of Prokaryotic names with Standing in Nomenclature (LPSN) is an online database that maintains information on the naming and taxonomy of prokaryotes, following the taxonomy requirements and rulings of the International Code of Nomenclature of Prokaryotes. The database was curated from 1997 to June 2013 by Jean P. Euzéby. From July 2013 to January 2020, LPSN was curated by Aidan C. Parte. In February 2020, a new version of LPSN was published as a service of the Leibniz Institute DSMZ, thereby also integrating the Prokaryotic Nomenclature Up-to-date service and since 2022 LPSN is interconnected with the Type (Strain) Genome Server (TYGS), DSMZ's high-throughput platform for accurate genome-based taxonomy. See also * Code of Nomenclature of Prokaryotes Described from Sequence Data References External links List of Prokaryotic names with Standing in Nomenclature
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Core Sample
A core sample is a cylindrical section of (usually) a naturally-occurring substance. Most core samples are obtained by drilling with special drills into the substance, such as sediment or rock, with a hollow steel tube, called a core drill. The hole made for the core sample is called the "core hole". A variety of core samplers exist to sample different media under different conditions; there is continuing development in the technology. In the coring process, the sample is pushed more or less intact into the tube. Removed from the tube in the laboratory, it is inspected and analyzed by different techniques and equipment depending on the type of data desired. Core samples can be taken to test the properties of manmade materials, such as concrete, ceramics, some metals and alloys, especially the softer ones. Core samples can also be taken of living things, including human beings, especially of a person's bones for microscopic examination to help diagnose diseases. Methods The ...
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