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Powdery Mildew (barley)
Powdery mildew is a fungal disease of barley caused by '' Blumeria graminis'' f. sp. ''hordei''. The disease has a worldwide distribution and is most damaging in cool, wet climates. The host range of the form species ''hordei'' is restricted to barley and other ''Hordeum species''. Symptoms At first, powdery mildew can be observed as small greyish patches of fluffy fungal growth (mycelium) on the upper surface of the lower leaves. These spots resemble small cushions of white powder. Leaf tissue on the opposite side of an infected leaf turns pale green to yellow. The fungus only infects the epidermal layer and can be easily scraped off with a fingernail. Infections can also occur on the leaf sheaths and ears. Leaves remain green and active for some time following infection, then gradually become chlorotic and die off. As the disease progresses, the mycelium often becomes dotted with minute black points ( cleistothecia), which are the sexual fruiting bodies of the fungus. ...
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Blumeria Graminis
''Blumeria graminis'' (commonly called barley powdery mildew, wheat mildew or corn mildew) is a fungus that causes powdery mildew on grasses, including cereals. It is the only species in the genus ''Blumeria''. It has also been called ''Erysiphe graminis'' and (by its anamorph) ''Oidium monilioides'' or ''Oidium tritici''. Systematics Previously ''B. graminis'' was included within the genus '' Erysiphe'', but molecular studies have placed it into a clade of its own. In 1975, it was moved to the new monospecific genus ''Blumeria''. ''Blumeria'' differs from ''Erysiphe'' in its digitate haustoria and in details of the conidial wall. ''Blumeria'' is also considered to be phylogenetically distinct from ''Erisiphe'' as it solely infects the true grasses of Poaceae. Eight special forms or '' formae speciales'' (ff.spp.) of ''B. graminis'' have been distinguished, each of which is parasitic on a particular genus or pareticular genera of grasses. Those that infect crop plants are ''B. ...
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Fungus
A fungus (: fungi , , , or ; or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and mold (fungus), molds, as well as the more familiar mushrooms. These organisms are classified as one of the kingdom (biology)#Six kingdoms (1998), traditional eukaryotic kingdoms, along with Animalia, Plantae, and either Protista or Protozoa and Chromista. A characteristic that places fungi in a different kingdom from plants, bacteria, and some protists is chitin in their cell walls. Fungi, like animals, are heterotrophs; they acquire their food by absorbing dissolved molecules, typically by secreting digestive enzymes into their environment. Fungi do not photosynthesize. Growth is their means of motility, mobility, except for spores (a few of which are flagellated), which may travel through the air or water. Fungi are the principal decomposers in ecological systems. These and other differences place fungi in a single group of related o ...
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Barley
Barley (), a member of the grass family, is a major cereal grain grown in temperate climates globally. It was one of the first cultivated grains; it was domesticated in the Fertile Crescent around 9000 BC, giving it nonshattering spikelets and making it much easier to harvest. Its use then spread throughout Eurasia by 2000 BC. Barley prefers relatively low temperatures and well-drained soil to grow. It is relatively tolerant of drought and soil salinity, but is less winter-hardy than wheat or rye. In 2023, barley was fourth among grains in quantity produced, 146 million tonnes, behind maize, rice, and wheat. Globally, 70% of barley production is used as animal feed, while 30% is used as a source of fermentable material for beer, or further distilled into whisky, and as a component of various foods. It is used in soups and stews and in barley bread of various cultures. Barley grains are commonly made into malt using a traditional and ancient method of preparatio ...
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Hordeum
''Hordeum'' is a genus of annual and perennial plants in the grass family. The species are native throughout the temperate regions of Africa, Eurasia, and the Americas. Taxonomy Species Species include: * '' Hordeum aegiceras'' – Mongolia, China including Tibet * '' Hordeum arizonicum'' US (CA AZ NV NM), Mexico (Baja California, Sonora, Durango) * '' Hordeum bogdanii'' – from Turkey and European Russia to Mongolia * '' Hordeum brachyantherum'' – Russia (Kuril, Kamchatka), Alaska, Canada including Yukon, US (mostly in the West but also scattered locales in the East), Baja California * '' Hordeum brachyatherum'' – Chile * '' Hordeum brevisubulatum'' – European Russia; temperate and subarctic Asia from Turkey and the Urals to China and Magadan * '' Hordeum bulbosum'' – Mediterranean, Central Asia * '' Hordeum californicum'' – US (CA; OR; NV) * '' Hordeum capense'' – South Africa, Lesotho * '' Hordeum chilense'' – Argentina, Chile (including Juan Fernández ...
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Mildew
Mildew is a form of fungus. It is distinguished from its closely related counterpart, mold, largely by its colour: molds appear in shades of black, blue, red, and green, whereas mildew is white. It appears as a thin, superficial growth consisting of minute hyphae (fungal filaments) produced especially on living plants or organic matter such as wood, paper or leather. Both mold and mildew produce distinct offensive odours, and both have been identified as the cause of certain human ailments. In horticulture, mildews are species of fungus in the order Erysiphales, or fungus-like organisms in the family '' Peronosporaceae''. It is also used more generally to mean mold growth. In Old English, mildew meant honeydew (a substance secreted by aphids on leaves, formerly thought to distill from the air like dew), and later came to mean mold or fungus. Household varieties The term mildew is often used generically to refer to mold growth, usually with a flat growth habit. Molds can ...
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Mycelium
Mycelium (: mycelia) is a root-like structure of a fungus consisting of a mass of branching, thread-like hyphae. Its normal form is that of branched, slender, entangled, anastomosing, hyaline threads. Fungal colonies composed of mycelium are found in and on soil and many other substrates. A typical single spore germinates into a monokaryotic mycelium, which cannot reproduce sexually; when two compatible monokaryotic mycelia join and form a dikaryotic mycelium, that mycelium may form fruiting bodies such as mushrooms. A mycelium may be minute, forming a colony that is too small to see, or may grow to span thousands of acres as in '' Armillaria''. Through the mycelium, a fungus absorbs nutrients from its environment. It does this in a two-stage process. First, the hyphae secrete enzymes onto or into the food source, which break down biological polymers into smaller units such as monomers. These monomers are then absorbed into the mycelium by facilitated diffusion and ac ...
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Chlorosis
In botany, chlorosis is a condition in which leaves produce insufficient chlorophyll. As chlorophyll is responsible for the green color of leaves, chlorotic leaves are pale, yellow, or yellow-white. The affected plant has little or no ability to manufacture carbohydrates through photosynthesis and may die unless the cause of its chlorophyll insufficiency is treated and this may lead to a plant disease called rusts, although some chlorotic plants, such as the albino ''Arabidopsis thaliana'' mutant ''ppi2'', are viable if supplied with exogenous sucrose. The word ''chlorosis'' is derived from the Greek ''khloros'' meaning "greenish-yellow", "pale green", "pale", "pallid", or "fresh". In viticulture, the most common symptom of poor nutrition in grapevines is the yellowing of grape leaves caused by chlorosis and the subsequent loss of chlorophyll. This is often seen in vineyard soils that are high in limestone such as the Italian wine region of Barolo in the Piedmont (wine), Piedmont ...
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Cleistothecia
An ascocarp, or ascoma (: ascomata), is the fruiting body ( sporocarp) of an ascomycete phylum fungus. It consists of very tightly interwoven hyphae and millions of embedded asci, each of which typically contains four to eight ascospores. Ascocarps are most commonly bowl-shaped (apothecia) but may take on a spherical or flask-like form that has a pore opening to release spores (perithecia) or no opening (cleistothecia). Classification The ascocarp is classified according to its placement (in ways not fundamental to the basic taxonomy). It is called ''epigeous'' if it grows above ground, as with the morels, while underground ascocarps, such as truffles, are termed ''hypogeous''. The structure enclosing the hymenium is divided into the types described below (apothecium, cleistothecium, etc.) and this character ''is'' important for the taxonomic classification of the fungus. Apothecia can be relatively large and fleshy, whereas the others are microscopic—about the size of ...
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Conidium
A conidium ( ; : conidia), sometimes termed an asexual chlamydospore or chlamydoconidium (: chlamydoconidia), is an Asexual reproduction, asexual, non-motility, motile spore of a fungus. The word ''conidium'' comes from the Ancient Greek word for dust, ('). They are also called mitospores due to the way they are generated through the cellular process of mitosis. They are produced exogenously. The two new haploid cells are genetically identical to the haploid parent, and can develop into new organisms if conditions are favorable, and serve in biological dispersal. Asexual reproduction in ascomycetes (the phylum Ascomycota) is by the formation of conidia, which are borne on specialized stalks called conidiophores. The Morphology (biology), morphology of these specialized conidiophores is often distinctive between species and, before the development of molecular techniques at the end of the 20th century, was widely used for identification of (''e.g.'' ''Metarhizium#Species, Metarh ...
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Ascospore
In fungi, an ascospore is the sexual spore formed inside an ascus—the sac-like cell that defines the division Ascomycota, the largest and most diverse Division (botany), division of fungi. After two parental cell nucleus, nuclei fuse, the ascus undergoes meiosis (halving of genetic material) followed by a mitosis (cell division), ordinarily producing eight genetically distinct haploid spores; most yeasts stop at four ascospores, whereas some moulds carry out extra post-meiotic divisions to yield dozens. Many asci build turgor, internal pressure and shoot their spores clear of the calm boundary layer, thin layer of still air enveloping the fruit body, whereas subterranean truffles depend on animals for biological dispersal, dispersal. Ontogeny, Development shapes both form and endurance of ascospores. A hook-shaped crozier aligns the paired nuclei; a double-biological membrane, membrane system then parcels each daughter nucleus, and successive wall layers of β-glucan, chitosan ...
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Propagule
In biology, a propagule is any material that functions in propagating an organism to the next stage in its life cycle, such as by dispersal. The propagule is usually distinct in form from the parent organism. Propagules are produced by organisms such as plants (in the form of seeds or spores), fungi (in the form of spores), and bacteria (for example endospores or microbial cysts). In disease biology, pathogens are said to generate infectious propagules, the units that transmit a disease. These can refer to bacteria, viruses, fungi, or protists, and can be contained within host material. For instance, for influenza, the infectious propagules are carried in droplets of host saliva or mucus that are expelled during coughing or sneezing. In horticulture, a propagule is any plant material used for the purpose of plant propagation. In asexual reproduction, a propagule is often a stem cutting. In some plants, a leaf section or a portion of root can be used. In sexual reprodu ...
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Sporulation
In biology, a spore is a unit of sexual (in fungi) or asexual reproduction that may be adapted for dispersal and for survival, often for extended periods of time, in unfavourable conditions. Spores form part of the life cycles of many plants, algae, fungi and protozoa. They were thought to have appeared as early as the mid-late Ordovician period as an adaptation of early land plants. Bacterial spores are not part of a sexual cycle, but are resistant structures used for survival under unfavourable conditions. Myxozoan spores release amoeboid infectious germs ("amoebulae") into their hosts for parasitic infection, but also reproduce within the hosts through the pairing of two nuclei within the plasmodium, which develops from the amoebula. In plants, spores are usually haploid and unicellular and are produced by meiosis in the sporangium of a diploid sporophyte. In some rare cases, a diploid spore is also produced in some algae, or fungi. Under favourable conditions, the s ...
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