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Portalia
''Portalia mira'' is an extinct species of uncertain placement, described by American paleontologist Charles Walcott from a fossil discovered in the Burgess Shale and initially interpreted as a holothurian echinoderm by Walcott, but this interpretation has been disputed. Taxonomy The taxonomy of this species remains unresolved. It was initially posited by Walcott that ''P. mira'' was a holothurian, but this was disputed by Frizzell and Exline (1955) and Madsen (1957), the latter of whom believed ''P. mira'' to be a kind of primitive sponge. J. Wyatt Durham wrote in 1974 that he felt that ''P. mira'' should not be rejected as a holothurian without further study. Description When examining the fossil holotype, the main body of ''P. mira'' is relatively long and contains a central strand that could represent the alimentary canal The gastrointestinal tract (GI tract, digestive tract, alimentary canal) is the tract or passageway of the digestive system that leads from the mouth ...
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Mackenzia
''Mackenzia'' is an elongated bag-like animal known from the Middle Cambrian Burgess Shale. It attached directly to hard surfaces, such as brachiopod shells. 14 specimens of ''Mackenzia'' are known from the Greater Phyllopod bed The Phyllopod bed, designated by USNM locality number 35k, is the most famous fossil-bearing member of the Burgess Shale fossil ''Lagerstätte''. It was quarried by Charles Walcott from 1911–1917 (and later named Walcott Quarry), and was ..., where they comprise <0.1% of the community. ''Mackenzia'' was originally described by Charles Walcott in 1911 as a holothurian echinoderm. Later, ''Mackenzia'' is thought to be a cnidarian and appears most similar to modern sea anemones.


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Charles Doolittle Walcott
Charles Doolittle Walcott (March 31, 1850February 9, 1927) was an American paleontologist, administrator of the Smithsonian Institution from 1907 to 1927, and director of the United States Geological Survey. He is famous for his discovery in 1909 of well-preserved fossils, including some of the oldest soft-part imprints, in the Burgess Shale of British Columbia, Canada. Early life Charles Doolittle Walcott was born on March 31, 1850, in New York Mills, New York. His grandfather, Benjamin S. Walcott, moved from Rhode Island in 1822. His father, also Charles Doolittle Walcott, died when Charles Jr. was only two. Walcott was the youngest of four children. He was interested in nature from an early age, collecting minerals and bird eggs and, eventually, fossils. He attended various schools in the Utica area but left at the age of eighteen without completing high school, the end of his formal education. His interest in fossils solidified as he became a commercial fossil collector ...
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Incertae Sedis
or is a term used for a taxonomy (biology), taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertainty at specific taxonomic levels is indicated by (of uncertain family), (of uncertain suborder), (of uncertain order) and similar terms. Examples * The fossil plant ''Paradinandra, Paradinandra suecica'' could not be assigned to any family, but was placed ''incertae sedis'' within the order Ericales when described in 2001. * The fossil ''Gluteus minimus (fossil), Gluteus minimus'', described in 1975, could not be assigned to any known animal phylum. The genus is therefore ''incertae sedis'' within the kingdom Animalia. * While it was unclear to which order the New World vultures (family Cathartidae) should be assigned, they were placed in Aves ''incertae sedis''. It was later agreed to place them in a separate order, Cathartiformes. * Boc ...
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Burgess Shale
The Burgess Shale is a fossil-bearing deposit exposed in the Canadian Rockies of British Columbia, Canada. It is famous for the exceptional preservation of the soft parts of its fossils. At old (middle Cambrian), it is one of the earliest fossil beds containing soft-part imprints. The rock unit is a black shale and crops out at a number of localities near the town of Field in Yoho National Park and the Kicking Horse Pass. Another outcrop is in Kootenay National Park 42 km to the south. History and significance The Burgess Shale was discovered by palaeontologist Charles Walcott on 30 August 1909, towards the end of the season's fieldwork. He returned in 1910 with his sons, daughter, and wife, establishing a quarry on the flanks of Fossil Ridge. The significance of soft-bodied preservation, and the range of organisms he recognised as new to science, led him to return to the quarry almost every year until 1924. At that point, aged 74, he had amassed over 65,000 s ...
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Holothurian
Sea cucumbers are echinoderms from the class Holothuroidea ( ). They are benthic marine animals found on the sea floor worldwide, and the number of known holothuroid species worldwide is about 1,786, with the greatest number being in the Asia–Pacific region. Sea cucumbers serve a useful role in the marine ecosystem as detritivores who help recycle nutrients, breaking down detritus and other organic matter, after which microbes can continue the decomposition process. Sea cucumbers have a leathery skin and an elongated body containing a single, branched gonad, are named for their overall resemblance to the fruit of the cucumber plant. Like all echinoderms, sea cucumbers have a calcified dermal endoskeleton, which is usually reduced to isolated microscopic ossicles (or sclerietes) joined by connective tissue. In some species these can sometimes be enlarged to flattened plates, forming an armoured cuticle. In some abyssal or pelagic species such as '' Pelagothuria natatrix'' (orde ...
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Echinoderm
An echinoderm () is any animal of the phylum Echinodermata (), which includes starfish, brittle stars, sea urchins, sand dollars and sea cucumbers, as well as the sessile sea lilies or "stone lilies". While bilaterally symmetrical as larvae, as adults echinoderms are recognisable by their usually five-pointed radial symmetry (pentamerous symmetry), and are found on the sea bed at every ocean depth from the intertidal zone to the abyssal zone. The phylum contains about 7,600 living species, making it the second-largest group of deuterostomes after the chordates, as well as the largest marine-only phylum. The first definitive echinoderms appeared near the start of the Cambrian. Echinoderms are important both ecologically and geologically. Ecologically, there are few other groupings so abundant in the deep sea, as well as shallower oceans. Most echinoderms are able to reproduce asexually and regenerate tissue, organs and limbs; in some cases, they can undergo ...
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Oxford University Press
Oxford University Press (OUP) is the publishing house of the University of Oxford. It is the largest university press in the world. Its first book was printed in Oxford in 1478, with the Press officially granted the legal right to print books by decree in 1586. It is the second-oldest university press after Cambridge University Press, which was founded in 1534. It is a department of the University of Oxford. It is governed by a group of 15 academics, the Delegates of the Press, appointed by the Vice Chancellor, vice-chancellor of the University of Oxford. The Delegates of the Press are led by the Secretary to the Delegates, who serves as OUP's chief executive and as its major representative on other university bodies. Oxford University Press has had a similar governance structure since the 17th century. The press is located on Walton Street, Oxford, Walton Street, Oxford, opposite Somerville College, Oxford, Somerville College, in the inner suburb of Jericho, Oxford, Jericho. ...
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Journal Of Paleontology
The ''Journal of Paleontology'' is a peer-reviewed scientific journal covering the field of paleontology. It is managed and published by Cambridge University Press on behalf of the Paleontological Society. Indexing The ''Journal of Paleontology'' is indexed in: *BIOSIS Previews *Science Citation Index *The Zoological Record *GeoRef __NOTOC__ The GeoRef database is a bibliographic database that indexes scientific literature in the geosciences, including geology. Coverage ranges from 1666 to the present for North American literature, and 1933 to the present for the rest of th ... References Paleontology journals Academic journals established in 1927 Academic journals published by learned and professional societies Cambridge University Press academic journals Bimonthly journals Paleontological Society {{paleo-journal-stub ...
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The Paleontological Society
The Paleontological Society, formerly the Paleontological Society of America, is an international organisation devoted to the promotion of paleontology. The Society was founded in 1908 in Baltimore, Maryland, and was incorporated in April 1968 in the District of Columbia. The Society publishes the bi-monthly ''Journal of Paleontology'' and the quarterly ''Paleobiology'', holds an annual meeting in the autumn in conjunction with the Geological Society of America, sponsors conferences and lectures, and provides grants and scholarships. The Society has five geographic sections—Pacific Coast (founded March 1911), North-Central (founded May 1974), Northeastern (founded March 1977), Southeastern (founded November 1979), Rocky Mountain (founded October 1985), and South-Central (founded November 1988). Medals and awards The Society recognizes distinguished accomplishments through three awards, one recognized by a medal, the other two by inscribed plaques normally presented annually: * ...
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Porifera
Sponges or sea sponges are primarily marine invertebrates of the animal phylum Porifera (; meaning 'pore bearer'), a Basal (phylogenetics) , basal clade and a sister taxon of the Eumetazoa , diploblasts. They are sessility (motility) , sessile filter feeders that are bound to the seabed, and are one of the most ancient members of macrobenthos, with many historical species being important sponge reef , reef-building organisms. Sponges are multicellular organisms consisting of jelly-like mesohyl sandwiched between two thin layers of cell (biology) , cells, and usually have tube-like bodies full of pores and channels that allow water to circulate through them. They have unspecialized cells that can cellular differentiation , transform into other types and that often migrate between the main cell layers and the mesohyl in the process. They do not have complex nervous system , nervous, digestive system , digestive or circulatory systems. Instead, most rely on maintaining a constant ...
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Type (biology)
In biology, a type is a particular specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally associated. In other words, a type is an example that serves to anchor or centralizes the defining features of that particular taxon. In older usage (pre-1900 in botany), a type was a taxon rather than a specimen. A taxon is a scientifically named grouping of organisms with other like organisms, a set that includes some organisms and excludes others, based on a detailed published description (for example a species description) and on the provision of type material, which is usually available to scientists for examination in a major museum research collection, or similar institution. Type specimen According to a precise set of rules laid down in the International Code of Zoological Nomenclature (ICZN) and the ''International Code of Nomenclature for algae, fungi, and plants'' (ICN), the scientific name of every taxon is ...
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Alimentary Canal
The gastrointestinal tract (GI tract, digestive tract, alimentary canal) is the tract or passageway of the digestive system that leads from the mouth to the anus. The tract is the largest of the body's systems, after the cardiovascular system. The GI tract contains all the major organs of the digestive system, in humans and other animals, including the esophagus, stomach, and intestines. Food taken in through the mouth is digested to extract nutrients and absorb energy, and the waste expelled at the anus as feces. ''Gastrointestinal'' is an adjective meaning of or pertaining to the stomach and intestines. Most animals have a "through-gut" or complete digestive tract. Exceptions are more primitive ones: sponges have small pores ( ostia) throughout their body for digestion and a larger dorsal pore ( osculum) for excretion, comb jellies have both a ventral mouth and dorsal anal pores, while cnidarians and acoels have a single pore for both digestion and excretion. The huma ...
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