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Pericycle
The pericycle is a cylinder of parenchyma or sclerenchyma cells that lies just inside the endodermis and is the outer most part of the stele of plants. Although it is composed of non-vascular parenchyma cells, it is still considered part of the vascular cylinder because it arises from the procambium as do the vascular tissues it surrounds. In eudicots, it also has the capacity to produce lateral roots. Branch roots arise from this primary meristem tissue. In plants undergoing secondary growth, the pericycle contributes to the vascular cambium often diverging into a cork cambium. In angiosperms certain molecules within the endodermis and the surrounding vasculature are sent to the pericycle which promotes the growth of the root meristems. Location The pericycle is located between the endodermis and phloem in plant roots. In dicot stems, it is situated around the ring of vascular bundles in the stele. Function In dicot roots, the pericycle strengthens the roots and provide ...
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Ground Tissue
The ground tissue of plants includes all tissues that are neither dermal nor vascular. It can be divided into three types based on the nature of the cell walls. This tissue system is present between the dermal tissue and forms the main bulk of the plant body. # Parenchyma cells have thin primary walls and usually remain alive after they become mature. Parenchyma forms the "filler" tissue in the soft parts of plants, and is usually present in cortex, pericycle, pith, and medullary rays in primary stem and root. # Collenchyma cells have thin primary walls with some areas of secondary thickening. Collenchyma provides extra mechanical and structural support, particularly in regions of new growth. # Sclerenchyma cells have thick lignified secondary walls and often die when mature. Sclerenchyma provides the main structural support to the plant. #Aerenchyma cells are found in aquatic plants. They are also known to be parenchyma cells with large air cavities surrounded by irregul ...
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Sclerenchyma
The ground tissue of plants includes all tissues that are neither dermal nor vascular. It can be divided into three types based on the nature of the cell walls. This tissue system is present between the dermal tissue and forms the main bulk of the plant body. # Parenchyma cells have thin primary walls and usually remain alive after they become mature. Parenchyma forms the "filler" tissue in the soft parts of plants, and is usually present in cortex, pericycle, pith, and medullary rays in primary stem and root. # Collenchyma cells have thin primary walls with some areas of secondary thickening. Collenchyma provides extra mechanical and structural support, particularly in regions of new growth. # Sclerenchyma cells have thick lignified secondary walls and often die when mature. Sclerenchyma provides the main structural support to the plant. #Aerenchyma cells are found in aquatic plants. They are also known to be parenchyma cells with large air cavities surrounded by irregular c ...
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Endodermis
The endodermis is the innermost layer of cortex in land plants. It is a cylinder of compact living cells, the radial walls of which are impregnated with hydrophobic substances ( Casparian strip) to restrict apoplastic flow of water to the inside. The endodermis is the boundary between the cortex and the stele. In many seedless plants, such as ferns, the endodermis is a distinct layer of cells immediately outside the vascular cylinder (stele) in roots and shoots. In most seed plants, especially woody types, the endodermis is present in roots but not in stems. The endodermis helps regulate the movement of water, ions and hormones into and out of the vascular system. It may also store starch, be involved in perception of gravity and protect the plant against toxins moving into the vascular system. Structure The endodermis is developmentally the innermost portion of the cortex. It may consist of a single layer of barrel-shaped cells without any intercellular spaces or sometimes s ...
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Stele (biology)
In a vascular plant, the stele is the central part of the root or Plant stem, stem containing the tissues derived from the procambium. These include vascular tissue, in some cases ground tissue (pith) and a pericycle, which, if present, defines the outermost boundary of the stele. Outside the stele lies the endodermis, which is the innermost cell layer of the cortex. The concept of the stele was developed in the late 19th century by France, French botanists Phillippe Édouard Léon van Tieghem, P. E. L. van Tieghem and H. Doultion as a model for understanding the relationship between the Shoot (botany), shoot and root, and for discussing the evolution of vascular plant morphology. Now, at the beginning of the 21st century, plant molecular biologists are coming to understand the genetics and developmental pathways that govern tissue patterns in the stele. Moreover, physiologists are examining how the anatomy (sizes and shapes) of different steles affect the function of organs. Proto ...
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Eudicots
The eudicots or eudicotyledons are flowering plants that have two seed leaves (cotyledons) upon germination. The term derives from ''dicotyledon'' (etymologically, ''eu'' = true; ''di'' = two; ''cotyledon'' = seed leaf). Historically, authors have used the terms tricolpates or non-magnoliid dicots. The current botanical terms were introduced in 1991, by evolutionary botanist James A. Doyle and paleobotanist Carol L. Hotton, to emphasize the later evolutionary divergence of tricolpate dicots from earlier, less specialized, dicots. Scores of familiar plants are eudicots, including many commonly cultivated and edible plants, numerous trees, tropicals and ornamentals. Among the most well-known eudicot genera are those of the sunflower (''Helianthus''), dandelion (''Taraxacum''), forget-me-not ('' Myosotis''), cabbage ('' Brassica''), apple (''Malus''), buttercup ('' Ranunculus''), maple ('' Acer'') and macadamia (''Macadamia''). Most leafy, mid-latitude trees are also classi ...
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Lateral Roots
Lateral roots, emerging from the pericycle (meristematic tissue), extend horizontally from the primary root (radicle) and over time makeup the iconic branching pattern of root systems. They contribute to anchoring the plant securely into the soil, increasing water uptake, and facilitate the extraction of nutrients required for the growth and development of the plant. Lateral roots increase the surface area of a plant's root system and can be found in great abundance in several plant species. In some cases, lateral roots have been found to form symbiotic relationships with rhizobia (bacteria) and mycorrhizae (fungi) found in the soil, to further increase surface area and increase nutrient uptake. Several factors are involved in the formation and development of lateral roots. Regulation of root formation is tightly controlled by plant hormones such as auxin, and by the precise control of aspects of the cell cycle. Such control can be particularly useful, as increased auxin levels he ...
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Meristem
In cell biology, the meristem is a structure composed of specialized tissue found in plants, consisting of stem cells, known as meristematic cells, which are undifferentiated cells capable of continuous cellular division. These meristematic cells play a fundamental role in plant growth, regeneration, and acclimatization, as they serve as the source of all differentiated plant tissues and organs. They contribute to the formation of structures such as fruits, leaves, and seeds, as well as supportive tissues like stems and roots. Meristematic cells are totipotent, meaning they have the ability to differentiate into any plant cell type. As they divide, they generate new cells, some of which remain meristematic cells while others differentiate into specialized cells that typically lose the ability to divide or produce new cell types. Due to their active division and undifferentiated nature, meristematic cells form the foundation for the formation of new plant organs and the c ...
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Secondary Growth
In botany, secondary growth is the growth that results from cell division in the cambia or lateral meristems and that causes the stems and roots to thicken, while primary growth is growth that occurs as a result of cell division at the tips of stems and roots, causing them to elongate, and gives rise to primary tissue. Secondary growth occurs in most seed plants, but monocots usually lack secondary growth. If they do have secondary growth, it differs from the typical pattern of other seed plants. The formation of secondary vascular tissues from the cambium is a characteristic feature of dicotyledons and gymnosperms. In certain monocots, the vascular tissues are also increased after the primary growth is completed but the cambium of these plants is of a different nature. In the living pteridophytes this feature is extremely rare, only occurring in '' Isoetes''. Lateral meristems In many vascular plants, secondary growth is the result of the activity of the two lateral me ...
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Vascular Cambium
The vascular cambium is the main growth tissue in the stems and roots of many plants exhibiting secondary growth, specifically in dicots such as buttercups and oak trees, gymnosperms such as pine trees, as well as in certain other vascular plants. It produces secondary xylem inwards, towards the pith, and secondary phloem outwards, towards the bark. Generally, more secondary xylem is produced than secondary phloem. In herbaceous plants, it occurs in the vascular bundles which are often arranged like beads on a necklace forming an interrupted ring inside the stem. In woody plants, it forms a cylinder of unspecialized meristem cells, as a continuous ring from which the new tissues are grown. Unlike the xylem and phloem, it does not transport water, minerals or food through the plant. Other names for the vascular cambium are the main cambium, wood cambium, or bifacial cambium. Occurrence Vascular cambia are found in all seed plants except for five angiosperm lineages whi ...
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Cork Cambium
Cork cambium (: cambia or cambiums) is a tissue found in many vascular plants as a part of the epidermis. It is one of the many layers of bark, between the cork and primary phloem. The cork cambium is a lateral meristem and is responsible for secondary growth that replaces the epidermis in roots and stems. It is found in woody and many herbaceous dicots, gymnosperms and some monocots (monocots usually lack secondary growth). It is one of the plant's meristems – the series of tissues consisting of embryonic disk (incompletely differentiated) cells from which the plant grows. The function of cork cambium is to produce the cork, a tough protective material.Junikka, L. (1994) "Macroscopic bark terminology". ''IAWA Journal'' 15(1): 3–45Trockenbrodt, M. (1990) "Survey and discussion of the terminology used in bark anatomy". ''IAWA Bulletin, New Series'' 11: 141–166 Synonyms for cork cambium are bark cambium, peri-cambium and phellogen. Phellogen is defined as the meristematic ...
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Angiosperms
Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed within a fruit. The group was formerly called Magnoliophyta. Angiosperms are by far the most diverse group of land plants with 64 orders, 416 families, approximately 13,000 known genera and 300,000 known species. They include all forbs (flowering plants without a woody stem), grasses and grass-like plants, a vast majority of broad-leaved trees, shrubs and vines, and most aquatic plants. Angiosperms are distinguished from the other major seed plant clade, the gymnosperms, by having flowers, xylem consisting of vessel elements instead of tracheids, endosperm within their seeds, and fruits that completely envelop the seeds. The ancestors of flowering plants diverged from the common ancestor of all living gymnosperms before the end of the Carbonifero ...
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Vascular Bundle
A vascular bundle is a part of the transport system in vascular plants. The transport itself happens in the stem, which exists in two forms: xylem and phloem. Both these tissues are present in a vascular bundle, which in addition will include supporting and protective tissues. There is also a tissue between xylem and phloem, which is the cambium. The xylem typically lies towards the axis ( adaxial) with phloem positioned away from the axis ( abaxial). In a stem or root this means that the xylem is closer to the centre of the stem or root while the phloem is closer to the exterior. In a leaf, the adaxial surface of the leaf will usually be the upper side, with the abaxial surface the lower side. The sugars synthesized by the plant with sun light are transported by the phloem, which is closer to the lower surface. Aphids and leaf hoppers feed off of these sugars by tapping into the phloem. This is why aphids and leaf hoppers are typically found on the underside of a l ...
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