Pachyosteus Bulla
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Pachyosteus Bulla
''Pachyosteus'' is an extinct monospecific genus of medium-sized selenosteid arthrodire placoderm known from the Upper Frasnian Kellwasserkalk facies of Late Devonian Bad Wildungen, Germany and from the Famennian portions of the Holy Cross Mountains of Poland. The type species ''Pachyosteus bulla'' has a broad skull about long, a comparatively long median dorsal plate, and a short rostral plate that meets the pineal plate. Phylogeny ''Pachyosteus'' is a member of the family Selenosteidae of the clade Aspinothoracidi, which belongs to the clade Pachyosteomorphi, one of the two major clades within Eubrachythoraci. The cladogram below shows the phylogeny of ''Pachyosteus'': In his cladogram, Rücklin (2011) regards ''Pachyosteus bulla'' as a basal selenosteid, being the sister taxon of the American genera, and the Kellwasserkalk genera of Germany and Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region o ...
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Late Devonian
The Devonian ( ) is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, million years ago (Mya), to the beginning of the Carboniferous, Mya. It is named after Devon, England, where rocks from this period were first studied. The first significant adaptive radiation of life on dry land occurred during the Devonian. Free-sporing vascular plants began to spread across dry land, forming extensive forests which covered the continents. By the middle of the Devonian, several groups of plants had evolved leaves and true roots, and by the end of the period the first seed-bearing plants appeared. The arthropod groups of myriapods, arachnids and hexapods also became well-established early in this period, after starting their expansion to land at least from the Ordovician period. Fish reached substantial diversity during this time, leading the Devonian to often be dubbed the Age of Fishes. The placoderms began dominating ...
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Poland
Poland, officially the Republic of Poland, , is a country in Central Europe. Poland is divided into Voivodeships of Poland, sixteen voivodeships and is the fifth most populous member state of the European Union (EU), with over 38 million people, and the List of European countries by area, seventh largest EU country, covering a combined area of . It extends from the Baltic Sea in the north to the Sudetes and Carpathian Mountains in the south, bordering seven countries. The territory is characterised by a varied landscape, diverse ecosystems, and Temperate climate, temperate transitional climate. The capital and List of cities and towns in Poland, largest city is Warsaw; other major cities include Kraków, Wrocław, Łódź, Poznań, and Gdańsk. Prehistory and protohistory of Poland, Humans have been present on Polish soil since the Lower Paleolithic, with continuous settlement since the end of the Last Glacial Period over 12,000 years ago. Culturally diverse throughout ...
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Eastmanosteus Calliaspis
''Eastmanosteus'' ("Eastman's bone") is a fossil genus of dunkleosteid placoderms. It was closely related to the giant ''Dunkleosteus'', but differed from that genus in size, in possessing a distinctive tuberculated bone ornament, a differently shaped nuchal plate and a more zig-zagging course of the sutures of the skull roof.http://www.ivpp.cas.cn/cbw/gjzdwxb/xbwzxz/200812/W020090813371138329343.pdf Species of ''Eastmanosteus'' had powerful jaws with sharp cutting edges and were likely active predators. Fossils have been found in many parts of the world in marine sediments dating from the Middle to Late Devonian. They were medium-to-large fish, with specimens ''E. pustulosus'' and ''E. licharevi'' approaching a total length of 3 metres. Complete exoskeletons with soft-tissue traces of ''E. calliaspis'' from Australia make this one of the best known dunkleosteids. Phylogeny ''Eastmanosteus'' and its relative ''Dunkleosteus'' belong to the family Dunkleosteidae. The phylogeny of ...
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Dunkleosteoidea
Dunkleosteoidea is an extinct superfamily of arthrodire placoderms that lived during the Devonian period. The gigantic apex predator '' Dunkleosteus terrelli'' is the best known member of this group. Phylogeny Eubrachythoraci is divided into the clades Coccosteomorphi and Pachyosteomorphi, the latter of which can be further sub-divided into Aspinothoracidi and Dunkleosteoidea. Dunkleosteoidea was then considered to consist of the two sister families Dunkleosteidae and Panxiosteidae. However, the 2016 Zhu ''et al.'' phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ... study using a larger morphological dataset recovered Panxiosteidae well outside of Dunkleosteoidea, leaving the status of Dunkleosteidae as a clade grouping separate from Dunkleosteoidea in doubt ...
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Rhachiosteus Pterygiatus
''Rhachiosteus'' is an extinct monospecific genus of arthrodire placoderm from the Middle to Late Devonian of Germany. It is known only from a single specimen, which may be a larval or juvenile form, as the skull of said specimen is only 19 millimetres long. Phylogeny ''Rhachiosteus'' is a basal member of the clade Pachyosteomorphi, the sister taxon to Coccosteomorphi, which together are the two main sub-clades of Eubrachythoraci. The cladogram below shows the phylogeny A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological spe ... of ''Rhachiosteus'': References Arthrodires Placoderms of Europe Arthrodire genera {{Placoderm-stub ...
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Coccosteomorphi
Coccosteomorphi is an extinct clade of arthrodire placoderms within the Eubrachythoraci (of the suborder Brachythoraci), armored fish most diverse during the Devonian. Most are considered to be pelagic (open ocean) long-distance swimmers, leading to their widespread distribution beginning from at least the Middle Devonian period. Phylogeny Coccosteomorphi is the sister taxon to Pachyosteomorphi, which together are the two main sub-clades of Eubrachythoraci. Coccosteomorphi can be further sub-divided into Coccosteoidea and Incisoscutoidea, as shown in the cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to ... below: References Arthrodires {{Placoderm-stub ...
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Zoological Journal Of The Linnean Society
The ''Zoological Journal of the Linnean Society'' is a monthly peer-reviewed scientific journal covering zoology published by Oxford University Press on behalf of the Linnean Society. The editor-in-chief is Maarten Christenhusz (Linnean Society). It was established in 1856 as the ''Journal of the Proceedings of the Linnean Society of London. Zoology'' and renamed ''Journal of the Linnean Society of London, Zoology'' in 1866. It obtained its current title in 1969. Abstracting and indexing The journal is abstracted and indexed in: According to the ''Journal Citation Reports'', the journal has a 2020 impact factor of 3.286. References External links

* Zoology journals Linnean Society of London Monthly journals Academic journals associated with learned and professional societies Publications established in 1856 {{zoo-journal-stub ...
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Phylogeny
A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological species or other entities based upon similarities and differences in their physical or genetic characteristics. All life on Earth is part of a single phylogenetic tree, indicating common ancestry. In a ''rooted'' phylogenetic tree, each node with descendants represents the inferred most recent common ancestor of those descendants, and the edge lengths in some trees may be interpreted as time estimates. Each node is called a taxonomic unit. Internal nodes are generally called hypothetical taxonomic units, as they cannot be directly observed. Trees are useful in fields of biology such as bioinformatics, systematics, and phylogenetics. ''Unrooted'' trees illustrate only the relatedness of the leaf nodes and do not require the ancestral root ...
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Cladogram
A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to descendants, nor does it show how much they have changed, so many differing evolutionary trees can be consistent with the same cladogram. A cladogram uses lines that branch off in different directions ending at a clade, a group of organisms with a last common ancestor. There are many shapes of cladograms but they all have lines that branch off from other lines. The lines can be traced back to where they branch off. These branching off points represent a hypothetical ancestor (not an actual entity) which can be inferred to exhibit the traits shared among the terminal taxa above it. This hypothetical ancestor might then provide clues about the order of evolution of various features, adaptation, and other evolutionary narratives about an ...
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Eubrachythoraci
Eubrachythoraci is an extinct clade of arthrodire placoderms within the suborder Brachythoraci, armored fish most diverse during the Devonian. Most are considered to be pelagic (open ocean) long-distance swimmers, leading to their widespread distribution beginning from at least the Middle Devonian period. Phylogeny Brachythoraci is divided into the large derived clade Eubrachythoraci and several basal groups: Buchanosteoidea, Homosteidae, and Holonematidae. (Although Holonematidae's membership in Brachythoraci is disputed.) Eubrachythoraci is then further divided into the sub-clades Coccosteomorphi and Pachyosteomorphi, the latter of which can be further sub-divided into Aspinothoracidi and Dunkleosteoidea, as shown in the cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are relate ...
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Pachyosteomorphi
Pachyosteomorphi is an extinct clade of arthrodire placoderms within the Eubrachythoraci (of the suborder Brachythoraci), armored fish most diverse during the Devonian. Most are considered to be pelagic (open ocean) long-distance swimmers, leading to their widespread distribution beginning from at least the Middle Devonian period. Phylogeny Pachyosteomorphi is the sister taxon to Coccosteomorphi, which together are the two main sub-clades of Eubrachythoraci. Pachyosteomorphi can be further sub-divided into Aspinothoracidi and Dunkleosteoidea, as shown in the cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to ... below: References Arthrodires {{Placoderm-stub ...
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Aspinothoracidi
Aspinothoracidi is a clade of placoderms, extinct armored fish most diverse during the Devonian. The gigantic apex predator '' Dinichthys'', is the best-known member of this group. Many other genera, such as the infamous '' Dunkleosteus'', were previously thought to be close relatives of ''Dinichthys'' and were grouped together in the family Dinichthyidae, though more recent studies have restricted that family to only its type species. Phylogeny Eubrachythoraci is divided into the clades Coccosteomorphi and Pachyosteomorphi, the latter of which can be further sub-divided into Aspinothoracidi and Dunkleosteoidea, as shown in the cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to ... below: References Arthrodires {{Placoderm-stub ...
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