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Pachycephalosaurini
Pachycephalosauria (; from Greek παχυκεφαλόσαυρος for 'thick headed lizards') is a clade of ornithischian dinosaurs. Along with Ceratopsia, it makes up the clade Marginocephalia. With the exception of two species, most pachycephalosaurs lived during the Late Cretaceous Period, dating between about 85.8 and 66 million years ago. They are exclusive to the Northern Hemisphere, all of them being found in North America and Asia. They were all bipedal, herbivorous/omnivorous animals with thick skulls. Skulls can be domed, flat, or wedge-shaped depending on the species, and are all heavily ossified. The domes were often surrounded by nodes and/or spikes. Partial skeletons have been found of several pachycephalosaur species, but to date no complete skeletons have been discovered. Often isolated skull fragments are the only bones that are found. Candidates for the earliest-known pachycephalosaur include ''Ferganocephale adenticulatum'' from Middle Jurassic Period strata o ...
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Alaskacephale
''Alaskacephale'' is an extinct genus of pachycephalosaurid, a group of dome-headed, herbivorous ornithischian dinosaurs, that lived during the Maastrichtian stage of the Late Cretaceous period in what is now northern Alaska. The genus is one of the few known Arctic dinosaurs and was found in the Prince Creek Formation, which preserves a menagerie of fossils. The only known specimen, a squamosal bone, was found in 1999 and later described in 2005. However, ''Alaskacephale'' was not formally named until the next year. ''Alaskacephale'', due to the lack of fossil remains, is poorly known. Despite this, the presence of bony protuberances from the skull prove that it was a pachycephalosaur. Later phylogenetic analyses suggest it was a close relative of ''Pachycephalosaurus''. However, ''Alaskacephale'' is distinguished from other genera by the two rows of nodules found along the squamosal. The Prince Creek Formation bears a variety of fossils, many of them similar to other previousl ...
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Pachycephalosaurus
''Pachycephalosaurus'' (; meaning "thick-headed lizard", from Greek ''pachys-/'' "thickness", ''kephalon/'' "head" and ''sauros/'' "lizard") is a genus of pachycephalosaurid ornithischian dinosaur. The type species, ''P. wyomingensis'', is the only known definitive species. The possibly synonymous taxon, ''Stygimoloch'', might represent a distinct genus or a second species, ''P. spinifer''. It lived during the Maastrichtian age of the Late Cretaceous period in what is now western North America. Remains have been excavated in Montana, South Dakota, Wyoming, and Alberta. Mainly known from a single skull and a few extremely thick skull roofs (at 22 cm or 9 in thick), ''Pachycephalosaurus'' is estimated to have reached long and weighed . More complete fossils would come to be found in the following years. ''Pachycephalosaurus'' was among the last species of non-avian dinosaurs on Earth before the Cretaceous–Paleogene extinction event. The genus ''Tylosteus'' has be ...
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Pachycephalosaurus Wyomingensis
''Pachycephalosaurus'' (; meaning "thick-headed lizard", from Ancient Greek, Greek ''pachys-/'' "thickness", ''kephalon/'' "head" and ''sauros/'' "lizard") is a genus of pachycephalosaurid Ornithischia, ornithischian dinosaur. The type species, ''P. wyomingensis'', is the only known definitive species. The possibly synonymous taxon, ''Stygimoloch'', might represent a distinct genus or a second species, ''P. spinifer''. It lived during the Maastrichtian Geologic time scale, age of the Late Cretaceous Period (geology), period in what is now Laramidia, western North America. Remains have been excavated in Montana, South Dakota, Wyoming, and Alberta. Mainly known from a single skull and a few extremely thick skull roofs (at 22 cm or 9 in thick), ''Pachycephalosaurus'' is estimated to have reached long and weighed . More complete fossils would come to be found in the following years. ''Pachycephalosaurus'' was among the last species of non-avian dinosaurs on Earth before ...
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Sphaerotholus
''Sphaerotholus'' is a genus of pachycephalosaurid dinosaur from the Upper Cretaceous of the western United States and Canada. To date, five species have been described: the type species, ''S. goodwini'', from the Den-na-zin Member of the Kirtland Formation and possibly the Fossil Forest Member of the Fruitland Formation (Late Campanian) of San Juan County, New Mexico, USA; ''S. buchholtzae'', from the Hell Creek Formation (Late Maastrichtian) of western Carter County, Montana, USA and the Frenchman Formation of Saskatchewan, Canada; ''S. edmontonensis'', from the Horseshoe Canyon Formation of Alberta, Canada; ''S. lyonsi'', from the Dinosaur Park Formation (Campanian) of Alberta, Canada; and ''S. triregnum'' from the Hell Creek Formation of Garfield County, Montana, USA. History of discovery The etymology of ''Sphaerotholus'' is a combination of the Greek ''sphaira'', meaning "ball", and ''tholos'', meaning "dome", and is a reference to the characteristically dome-shaped pa ...
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Texacephale
''Texacephale'' is a possibly dubious genus of pachycephalosaurid dinosaur from the Campanian stage of the Late Cretaceous. Its fossils come from the Aguja Formation of Big Bend National Park, in Texas, and were described in 2010 by Longrich, Sankey and Tanke. The generic name means Texas + "head" (''kephale'' in Greek) in reference to its place of discovery, and the specific name ''langstoni'' honors Wann Langston. It may be a synonym of ''Stegoceras.'' Discovery The holotype specimen of ''Texacephale'', LSUMNS 20010, is composed of fused frontals and parietals. A second specimen, LSUMNS 20012, is composed of an incomplete frontoparietal dome. It was found in the same WPA quarries that produced '' Agujaceratops'', and may have been excavated and tossed aside, being mistaken for a rock or concretion, before being picked up decades later. According to the team, the fossilized dome of the animal possessed five to six vertical flanges on each lateral side, connecting it with t ...
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Ornithischian
Ornithischia () is an extinct clade of mainly herbivorous dinosaurs characterized by a pelvic structure superficially similar to that of birds. The name ''Ornithischia'', or "bird-hipped", reflects this similarity and is derived from the Greek stem ' (), meaning "bird", and ' (), meaning "hip". However, as theropod dinosaurs, birds are only distantly related to this group. Ornithischians with well known anatomical adaptations include the ceratopsians or "horn-faced" dinosaurs (e.g. ''Triceratops''), the pachycephalosaurs or "thick-headed" dinosaurs, the armored dinosaurs (Thyreophora) such as stegosaurs and ankylosaurs, and the ornithopods. There is strong evidence that certain groups of ornithischians lived in herds, often segregated by age group, with juveniles forming their own flocks separate from adults. Some were at least partially covered in filamentous (hair- or feather- like) pelts, and there is much debate over whether these filaments found in specimens of '' Tianyulong ...
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Stegoceras
''Stegoceras'' is a genus of Pachycephalosauria, pachycephalosaurid (dome-headed) dinosaur that lived in what is now North America during the Late Cretaceous Period (geology), period, about 77.5 to 74 million years ago (mya). The first specimens from Alberta, Canada, were described in 1902, and the type species ''Stegoceras validum'' was based on these remains. The generic name means "horn roof", and the specific name means "strong". Several other species have been placed in the genus over the years, but these have since been moved to other genera or deemed junior synonyms. Currently only ''S. validum'' and ''S. novomexicanum'', named in 2011 from fossils found in New Mexico, remain. The validity of the latter species has also been debated, and it may not even belong to the genus ''Stegoceras.'' ''Stegoceras'' was a small, bipedal dinosaur about long, and weighed around . The skull was roughly triangular with a short snout, and had a thick, broad, and relatively smooth dome o ...
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Platytholus
''Platytholus'' is an extinct genus of pachycephalosaurid dinosaur from the Late Cretaceous (Maastrichtian) Hell Creek Formation of the United States. The genus contains a single species, ''P. clemensi'', known from a partial skull. Discovery and naming The ''Platytholus'' holotype specimen, MOR 2915, was discovered during the early 2000s in sediments of the Hell Creek Formation, dated to the latest Maastrichtian age of the late Cretaceous period, in Garfield County, Montana. The specimen consists of a partial skull. In 2023, Horner, Goodwin & Evans described ''Platytholus clemensi'', a new genus and species of pachycephalosaurid, based on these fossil remains. The generic name, "''Platytholus''", combines the Latinised Greek "platys", meaning "flattened", "wide", or "broad", and "tholos", meaning a "small domed hill". The specific name, "''clemensi''", honors American paleontologist William A. Clemens Jr. Classification Horner, Goodwin & Evans (2023) recovered ''Pla ...
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Sinocephale
''Sinocephale'' (meaning "Chinese head") is a genus of pachycephalosaurid dinosaur that lived in Inner Mongolia, China during the Cretaceous period. The only species, ''Sinocephale bexelli'', was originally named as a species of the genus ''Troodon'' in 1953, and later transferred to the genus ''Stegoceras''. After decades of being considered nomen dubium, dubious, it was re-evaluated in 2021 and recognized as a valid taxon, being given a unique generic name. The original holotype was lost, with modern research conducted using rediscovered plaster casts. Scant material makes for limited knowledge of its life appearance, but it is distinguished by an embayment on the back of the domed skull, which would give it a heart shape as seen from above. It is potentially the oldest known pachycephalosaurid and falls within the subset of the family called Pachycephalosaurinae, related to animals such as ''Pachycephalosaurus'' and ''Prenocephale''. The geologic context of the species has been ...
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Tylocephale
''Tylocephale'' (meaning "swollen head") is a genus of pachycephalosaurid dinosaur, a group of dome-headed, herbivorous ornithischians, that lived during the Late Campanian stage (75-73 million years ago) of the Late Cretaceous in what is now Mongolia. It is known from a partial skull and associated mandible that were unearthed in 1971 by a Polish-Mongolian Expedition to the Barun Goyot Formation of the Gobi Desert. The specimen was described in 1974 by Polish paleontologists Teresa Maryańska and Halszka Osmólska as a new genus and species. It was average-sized for a pachycephalosaur, reaching in length and in body mass. The skull is triangular in back view, the widest point being at the jugals with an apex at the top of the dome. ''Tylocephale'''s dome is the tallest known from a pachycephalosaur. This dome is also unusually thick and rugose on its exterior. Behind the dome, an array of spikes, nodes, and tubercules protrude posteriorly over the neck. The eyebrow ridge wa ...
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Oxford University Press
Oxford University Press (OUP) is the publishing house of the University of Oxford. It is the largest university press in the world. Its first book was printed in Oxford in 1478, with the Press officially granted the legal right to print books by decree in 1586. It is the second-oldest university press after Cambridge University Press, which was founded in 1534. It is a department of the University of Oxford. It is governed by a group of 15 academics, the Delegates of the Press, appointed by the Vice Chancellor, vice-chancellor of the University of Oxford. The Delegates of the Press are led by the Secretary to the Delegates, who serves as OUP's chief executive and as its major representative on other university bodies. Oxford University Press has had a similar governance structure since the 17th century. The press is located on Walton Street, Oxford, Walton Street, Oxford, opposite Somerville College, Oxford, Somerville College, in the inner suburb of Jericho, Oxford, Jericho. ...
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Lexico
''Lexico'' was a dictionary website that provided a collection of English and Spanish dictionaries produced by Oxford University Press (OUP), the publishing house of the University of Oxford. While the dictionary content on ''Lexico'' came from OUP, this website was operated by Dictionary.com, whose eponymous website hosts dictionaries by other publishers such as Random House. The website was closed and redirected to Dictionary.com on 26 August 2022. Before the Lexico site was launched, the ''Oxford Dictionary of English'' and ''New Oxford American Dictionary'' were hosted by OUP's own website ''Oxford Dictionaries Online'' (''ODO''), later known as ''Oxford Living Dictionaries''. The dictionaries' definitions have also appeared in Google Dictionary, Google definition search and the Dictionary (software), Dictionary application on macOS, among others, licensed through the Oxford Dictionaries API. History In the 2000s, OUP allowed access to content of the ''Compact Oxford Englis ...
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