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Pachycephalosauridae
Pachycephalosauria (; from Greek παχυκεφαλόσαυρος for 'thick headed lizards') is a clade of ornithischian dinosaurs. Along with Ceratopsia, it makes up the clade Marginocephalia. With the exception of two species, most pachycephalosaurs lived during the Late Cretaceous Period, dating between about 85.8 and 66 million years ago. They are exclusive to the Northern Hemisphere, all of them being found in North America and Asia. They were all bipedal, herbivorous/omnivorous animals with thick skulls. Skulls can be domed, flat, or wedge-shaped depending on the species, and are all heavily ossified. The domes were often surrounded by nodes and/or spikes. Partial skeletons have been found of several pachycephalosaur species, but to date no complete skeletons have been discovered. Often isolated skull fragments are the only bones that are found. Candidates for the earliest-known pachycephalosaur include ''Ferganocephale adenticulatum'' from Middle Jurassic Period strata ...
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Tylocephale
''Tylocephale'' (meaning "swollen head") is a genus of pachycephalosaurid dinosaur, a group of dome-headed, herbivorous ornithischians, that lived during the Late Campanian stage (75-73 million years ago) of the Late Cretaceous in what is now Mongolia. It is known from a partial skull and associated mandible that were unearthed in 1971 by a Polish-Mongolian Expedition to the Barun Goyot Formation of the Gobi Desert. The specimen was described in 1974 by Polish paleontologists Teresa Maryańska and Halszka Osmólska as a new genus and species. It was average-sized for a pachycephalosaur, reaching in length and in body mass. The skull is triangular in back view, the widest point being at the jugals with an apex at the top of the dome. ''Tylocephale'''s dome is the tallest known from a pachycephalosaur. This dome is also unusually thick and rugose on its exterior. Behind the dome, an array of spikes, nodes, and tubercules protrude posteriorly over the neck. The eyebrow ridge wa ...
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Goyocephale
''Goyocephale'' is an extinct genus of pachycephalosaurian ornithischian that lived in Mongolia during the Late Cretaceous about 76 million years ago. It was first described in 1982 by Altangerel Perle, Teresa Maryańska and Halszka Osmólska for a disarticulated skeleton with most of a skull, part of the forelimb and hindlimb, some of the pelvic girdle, and some vertebrae. Perle ''et al.'' named the remains ''Goyocephale lattimorei'', from the Mongolian language, Mongolian гоё (''goyo''), meaning "decorated", and the Ancient Greek κεφαλή (''kephale''), for head. The species name honours Owen Lattimore. Description ''Goyocephale'' is known from a partial skull, including both mandibles, the skull roof, part of the occiput, part of the braincase region, the anatomical terms of location#Anterior and posterior, posterior skull, the premaxilla, and the maxilla. The posterior edge of the skull roof, at the edge of the squamosal bones, has many small bony bumps, which would ...
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Texacephale
''Texacephale'' is a possibly dubious genus of pachycephalosaurid dinosaur from the Campanian stage of the Late Cretaceous. Its fossils come from the Aguja Formation of Big Bend National Park, in Texas, and were described in 2010 by Longrich, Sankey and Tanke. The generic name means Texas + "head" (''kephale'' in Greek) in reference to its place of discovery, and the specific name ''langstoni'' honors Wann Langston. It may be a synonym of ''Stegoceras.'' Discovery The holotype specimen of ''Texacephale'', LSUMNS 20010, is composed of fused frontals and parietals. A second specimen, LSUMNS 20012, is composed of an incomplete frontoparietal dome. It was found in the same WPA quarries that produced '' Agujaceratops'', and may have been excavated and tossed aside, being mistaken for a rock or concretion, before being picked up decades later. According to the team, the fossilized dome of the animal possessed five to six vertical flanges on each lateral side, connecting it with t ...
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Stegoceras
''Stegoceras'' is a genus of Pachycephalosauria, pachycephalosaurid (dome-headed) dinosaur that lived in what is now North America during the Late Cretaceous Period (geology), period, about 77.5 to 74 million years ago (mya). The first specimens from Alberta, Canada, were described in 1902, and the type species ''Stegoceras validum'' was based on these remains. The generic name means "horn roof", and the specific name means "strong". Several other species have been placed in the genus over the years, but these have since been moved to other genera or deemed junior synonyms. Currently only ''S. validum'' and ''S. novomexicanum'', named in 2011 from fossils found in New Mexico, remain. The validity of the latter species has also been debated, and it may not even belong to the genus ''Stegoceras.'' ''Stegoceras'' was a small, bipedal dinosaur about long, and weighed around . The skull was roughly triangular with a short snout, and had a thick, broad, and relatively smooth dome o ...
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Amtocephale
''Amtocephale'' is a genus of pachycephalosaurid dinosaur from early Late Cretaceous (Cenomanian-Coniacian stages) deposits of southern Gobi Desert, Mongolia. ''Amtocephale'' is known from the holotype MPC-D 100/1203, a nearly complete frontoparietal dome of a subadult individual. It was collected from the Baynshire Formation at the Amtgai locality. ''Amtocephale'' was first named by Mahito Watabe, Khishigjaw Tsogtbaatar and Robert M. Sullivan in 2011 and the type species is ''Amtocephale gobiensis''. The generic name combines a reference to the Amtgai site with a Greek κεφαλή, ''kephale'', "head". The specific name refers to the provenance from the Gobi. ''Amtocephale'' was assigned to the Pachycephalosauridae and is perhaps the oldest pachycephalosaurid known, depending on the exact age of the formation. History of naming Paleontological expeditions of the Hayashibara Museum of Natural Sciences with the Mongolian Paleontological Center into the Gobi Desert bega ...
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Dinosaur
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutionary history, evolution of dinosaurs is a subject of active research. They became the dominant terrestrial vertebrates after the Triassic–Jurassic extinction event 201.3 mya and their dominance continued throughout the Jurassic and Cretaceous periods. The fossil record shows that birds are feathered dinosaurs, Evolution of birds, having evolved from earlier Theropoda, theropods during the Late Jurassic epoch, and are the only dinosaur lineage known to have survived the Cretaceous–Paleogene extinction event approximately 66 mya. Dinosaurs can therefore be divided into avian dinosaurs—birds—and the extinct non-avian dinosaurs, which are all dinosaurs other than birds. Dinosaurs are varied from taxonomy (biology), taxonomic, ...
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Ornithischian
Ornithischia () is an extinct clade of mainly herbivorous dinosaurs characterized by a pelvic structure superficially similar to that of birds. The name ''Ornithischia'', or "bird-hipped", reflects this similarity and is derived from the Greek stem ' (), meaning "bird", and ' (), meaning "hip". However, as theropod dinosaurs, birds are only distantly related to this group. Ornithischians with well known anatomical adaptations include the ceratopsians or "horn-faced" dinosaurs (e.g. ''Triceratops''), the pachycephalosaurs or "thick-headed" dinosaurs, the armored dinosaurs (Thyreophora) such as stegosaurs and ankylosaurs, and the ornithopods. There is strong evidence that certain groups of ornithischians lived in herds, often segregated by age group, with juveniles forming their own flocks separate from adults. Some were at least partially covered in filamentous (hair- or feather- like) pelts, and there is much debate over whether these filaments found in specimens of '' Tianyulong ...
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Clade
In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach to taxonomy adopted by most biological fields. The common ancestor may be an individual, a population, or a species (extinct or Extant taxon, extant). Clades are nested, one in another, as each branch in turn splits into smaller branches. These splits reflect evolutionary history as populations diverged and evolved independently. Clades are termed ''monophyletic'' (Greek: "one clan") groups. Over the last few decades, the cladistic approach has revolutionized biological classification and revealed surprising evolutionary relationships among organisms. Increasingly, taxonomists try to avoid naming Taxon, taxa that are not clades; that is, taxa that are not Monophyly, monophyletic. Some of the relationships between organisms that the molecul ...
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Ancient Greek
Ancient Greek (, ; ) includes the forms of the Greek language used in ancient Greece and the classical antiquity, ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Greek Dark Ages, Dark Ages (), the Archaic Greece, Archaic or Homeric Greek, Homeric period (), and the Classical Greece, Classical period (). Ancient Greek was the language of Homer and of fifth-century Athens, fifth-century Athenian historians, playwrights, and Ancient Greek philosophy, philosophers. It has contributed many words to English vocabulary and has been a standard subject of study in educational institutions of the Western world since the Renaissance. This article primarily contains information about the Homeric Greek, Epic and Classical periods of the language, which are the best-attested periods and considered most typical of Ancient Greek. From the Hellenistic period (), Ancient Greek was followed by Koine Greek, which is regar ...
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Oxford University Press
Oxford University Press (OUP) is the publishing house of the University of Oxford. It is the largest university press in the world. Its first book was printed in Oxford in 1478, with the Press officially granted the legal right to print books by decree in 1586. It is the second-oldest university press after Cambridge University Press, which was founded in 1534. It is a department of the University of Oxford. It is governed by a group of 15 academics, the Delegates of the Press, appointed by the Vice Chancellor, vice-chancellor of the University of Oxford. The Delegates of the Press are led by the Secretary to the Delegates, who serves as OUP's chief executive and as its major representative on other university bodies. Oxford University Press has had a similar governance structure since the 17th century. The press is located on Walton Street, Oxford, Walton Street, Oxford, opposite Somerville College, Oxford, Somerville College, in the inner suburb of Jericho, Oxford, Jericho. ...
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Lexico
''Lexico'' was a dictionary website that provided a collection of English and Spanish dictionaries produced by Oxford University Press (OUP), the publishing house of the University of Oxford. While the dictionary content on ''Lexico'' came from OUP, this website was operated by Dictionary.com, whose eponymous website hosts dictionaries by other publishers such as Random House. The website was closed and redirected to Dictionary.com on 26 August 2022. Before the Lexico site was launched, the ''Oxford Dictionary of English'' and ''New Oxford American Dictionary'' were hosted by OUP's own website ''Oxford Dictionaries Online'' (''ODO''), later known as ''Oxford Living Dictionaries''. The dictionaries' definitions have also appeared in Google Dictionary, Google definition search and the Dictionary (software), Dictionary application on macOS, among others, licensed through the Oxford Dictionaries API. History In the 2000s, OUP allowed access to content of the ''Compact Oxford Englis ...
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Alaskacephale
''Alaskacephale'' is an extinct genus of pachycephalosaurid, a group of dome-headed, herbivorous ornithischian dinosaurs, that lived during the Maastrichtian stage of the Late Cretaceous period in what is now northern Alaska. The genus is one of the few known Arctic dinosaurs and was found in the Prince Creek Formation, which preserves a menagerie of fossils. The only known specimen, a squamosal bone, was found in 1999 and later described in 2005. However, ''Alaskacephale'' was not formally named until the next year. ''Alaskacephale'', due to the lack of fossil remains, is poorly known. Despite this, the presence of bony protuberances from the skull prove that it was a pachycephalosaur. Later phylogenetic analyses suggest it was a close relative of ''Pachycephalosaurus''. However, ''Alaskacephale'' is distinguished from other genera by the two rows of nodules found along the squamosal. The Prince Creek Formation bears a variety of fossils, many of them similar to other previousl ...
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