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Oxymonadida
The Oxymonads (or Oxymonadida) are a group of flagellated protists found exclusively in the intestines of animals, mostly termites and other wood-eating insects. Along with the similar parabasalid flagellates, they harbor the symbiotic bacteria that are responsible for breaking down cellulose. There is no evidence for presence of mitochondria (not even anaerobic mitochondrion-like organelles like hydrogenosomes or mitosomes) in oxymonads and three species have been shown to completely lack any molecular markers of mitochondria. It includes e.g. '' Dinenympha'', '' Pyrsonympha'', '' Oxymonas'', ''Streblomastix'', ''Monocercomonoides'', and '' Blattamonas''. Characteristics Most Oxymonads are around 50 μm in size and have a single nucleus, associated with four flagella. Their basal bodies give rise to several long sheets of microtubules, which form an organelle called an axostyle, but different in structure from the axostyles of parabasalids. The cell may use the axostyle to s ...
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Oxymonas
''Oxymonas'' is a genus of Excavata. History The genus ''Oxymonas'' within Excavata Excavata is an obsolete, extensive and diverse paraphyletic group of unicellular Eukaryota. The group was first suggested by Simpson and Patterson in 1999 and the name latinized and assigned a rank by Thomas Cavalier-Smith in 2002. It contains ..., was discovered by Janicki in 1915 within termites. ''Oxymonas'' was established in order to accommodate a newly discovered species, ''Oxymonas granulosa''. Through the discovery of ''Oxymonas'', Janicki was also responsible for coining the term karyomastigont for a new structure which was observed in this genus and other termite symbionts. Structure ''Oxymonas'' is found to be sub-elliptical in its body shape and has a pointed posterior end. ''Oxymonas'' is distinguishable by the rostellum, which can be thought of as an elongated proboscis. The rostellum projects anteriorly from the organism and ends in a holdfast apparatus that allows for the att ...
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Monocercomonoides
''Monocercomonoides'' is a genus of flagellate Excavata belonging to the order Oxymonadida. It was established by Bernard V. Travis and was first described as those with "polymastiginid flagellates having three anterior Flagellum, flagella and a trailing one originating at a single basal granule located in front of the anteriorly positioned cell nucleus, nucleus, and a more or less well-defined axostyle". It is the first eukaryotic genus to be found to completely lack Mitochondrion, mitochondria, and all hallmark proteins responsible for mitochondrial function. The genus also lacks any other Mitochondrion#Origin and evolution, mitochondria related organelles (MROs) such as hydrogenosomes or mitosomes. Data suggests that the absence of mitochondria is not an ancestral feature, but rather due to secondary loss. ''Monocercomonoides ''sp. was found to obtain energy through an enzymatic action of nutrients absorbed from the environment. The genus has replaced the Iron–sulfur protein ...
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Preaxostyla
Anaeromonadea, also known as Preaxostyla, is a class of excavate protists, comprising the oxymonads, '' Trimastix'', and '' Paratrimastix''. This group is studied as a model system for reductive evolution of mitochondria, because it includes both organisms with anaerobic mitochondrion-like organelles ('' Trimastix'' and '' Paratrimastix''), and those that have completely lost their mitochondria ( oxymonads ''Monocercomonoides'', ''Streblomastix A symbiotic eukaryote that lives in the hindgut of termites, ''Streblomastix'' is a protist associated with a community of ectosymbiotic bacteria. Motility ''Streblomastix'' moves by beating its anterior flagella. Morphology These protists ...'', and '' Blattamonas''). Phylogeny and Taxonomy Based on the work of Zhang et al. 2015. References External links Tree of Life: Preaxostyla Metamonads {{Excavata-stub ...
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Pyrsonympha
''Pyrsonympha'' is a genus of Excavata. It includes the species ''Pyrsonympha vertens''. Species List of ''Pyrsonympha'' species. * ''P. affinis'' Fedorowa 1923 * ''P. elongata'' Georgevitch 1932 * ''P. flagellata'' Grassé 1894 * ''P. grandis'' Koidzumi 1921 * ''P. granulata'' Powell 1928 * ''P. havilalldi'' Das 1972 * ''P. major'' Powell 1928 * ''P. minor'' Powell 1928 * ''P. modesta'' Koidzumi 1921 * ''P. omblensis'' Georgevitch 1951 * ''P. rostrata'' Mukherjee & Maiti 1988 * ''P. tirapi'' Mukherjee & Maiti 1988 * ''P. vertens'' Leidy 1877 References Metamonads Metamonad genera {{Excavata-stub ...
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Dinenympha
''Dinenympha'' is a genus of Excavata Excavata is an obsolete, extensive and diverse paraphyletic group of unicellular Eukaryota. The group was first suggested by Simpson and Patterson in 1999 and the name latinized and assigned a rank by Thomas Cavalier-Smith in 2002. It contains .... It includes the species ''Dinenympha exilis''. Species List of ''Dinenympha'' species. * ''D. aculeata'' Georgevitch 1951 * ''D. aviformis'' Georgevitch 1951 * ''D. exilis'' Koidzumi 1921 * ''D. fimbriata'' Kirby 1924 * ''D. gracilis'' Leidy 1877 * ''D. leidyi'' Koidzumi 1921 * ''D. mukundai'' Mukherjee & Maiti 1989 * ''D. nobilis'' Koidzumi 1921 * ''D. parva'' Koidzumi 1921 * ''D. porteri'' Koidzumi 1921 * ''D. rayi'' Mukherjee & Maiti 1989 * ''D. rugosa'' Koidzumi 1921 References Metamonad genera Metamonads {{Excavata-stub ...
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Cellulose
Cellulose is an organic compound with the chemical formula, formula , a polysaccharide consisting of a linear chain of several hundred to many thousands of glycosidic bond, β(1→4) linked glucose, D-glucose units. Cellulose is an important structural component of the primary cell wall of green plants, many forms of algae and the oomycetes. Some species of bacteria secrete it to form biofilms. Cellulose is the most abundant biopolymer, organic polymer on Earth. The cellulose content of cotton fibre is 90%, that of wood is 40–50%, and that of dried hemp is approximately 57%. Cellulose is mainly used to produce paperboard and paper. Smaller quantities are converted into a wide variety of derivative products such as cellophane and rayon. Conversion of cellulose from energy crops into biofuels such as cellulosic ethanol is under development as a renewable fuel source. Cellulose for industrial use is mainly obtained from wood pulp and cotton. Cellulose is also greatly affected by ...
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Axostyle
An axostyle is a sheet of microtubules found in certain protists. It arises from the bases of the flagella, sometimes projecting beyond the end of the cell, and is often flexible or contractile, and so may be involved in movement and provides support for the cell. Axostyles originate in association with a flagellar microtubular root and occur in two groups, the oxymonads and parabasalids; they have different structures and are not homologous. Within trichomonads the axostyle has been theorised to participate in locomotion and cell adhesion, but also karyokinesis Mitosis () is a part of the cell cycle in eukaryotic cells in which replicated chromosomes are separated into two new nuclei. Cell division by mitosis is an equational division which gives rise to genetically identical cells in which the ... during cell division. References {{Protist Cell biology ...
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Microtubule
Microtubules are polymers of tubulin that form part of the cytoskeleton and provide structure and shape to eukaryotic cells. Microtubules can be as long as 50 micrometres, as wide as 23 to 27 nanometer, nm and have an inner diameter between 11 and 15 nm. They are formed by the polymerization of a Protein dimer, dimer of two globular proteins, Tubulin#Eukaryotic, alpha and beta tubulin into #Structure, protofilaments that can then associate laterally to form a hollow tube, the microtubule. The most common form of a microtubule consists of 13 protofilaments in the tubular arrangement. Microtubules play an important role in a number of cellular processes. They are involved in maintaining the structure of the cell and, together with microfilaments and intermediate filaments, they form the cytoskeleton. They also make up the internal structure of cilia and flagella. They provide platforms for intracellular transport and are involved in a variety of cellular processes, in ...
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Basal Body
A basal body (synonymous with basal granule, kinetosome, and in older cytological literature with blepharoplast) is a protein structure found at the base of a eukaryotic undulipodium (cilium or flagellum). The basal body was named by Theodor Wilhelm Engelmann in 1880. It is formed from a centriole and several additional protein structures, and is, essentially, a modified centriole. The basal body serves as a nucleation site for the growth of the axoneme microtubules. Centrioles, from which basal bodies are derived, act as anchoring sites for proteins that in turn anchor microtubules, and are known as the microtubule organizing center (MTOC). These microtubules provide structure and facilitate movement of vesicles and organelles within many eukaryotic cells. Assembly, structure Cilia and basal bodies form during quiescence or the G1 phase of the cell cycle. Before the cell enters G1 phase, i.e. before the formation of the cilium, the mother centriole serves as a component of ...
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Flagellum
A flagellum (; : flagella) (Latin for 'whip' or 'scourge') is a hair-like appendage that protrudes from certain plant and animal sperm cells, from fungal spores ( zoospores), and from a wide range of microorganisms to provide motility. Many protists with flagella are known as flagellates. A microorganism may have from one to many flagella. A gram-negative bacterium '' Helicobacter pylori'', for example, uses its flagella to propel itself through the stomach to reach the mucous lining where it may colonise the epithelium and potentially cause gastritis, and ulcers – a risk factor for stomach cancer. In some swarming bacteria, the flagellum can also function as a sensory organelle, being sensitive to wetness outside the cell. Across the three domains of Bacteria, Archaea, and Eukaryota, the flagellum has a different structure, protein composition, and mechanism of propulsion but shares the same function of providing motility. The Latin word means " whip" to describe its ...
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Cell Nucleus
The cell nucleus (; : nuclei) is a membrane-bound organelle found in eukaryote, eukaryotic cell (biology), cells. Eukaryotic cells usually have a single nucleus, but a few cell types, such as mammalian red blood cells, have #Anucleated_cells, no nuclei, and a few others including osteoclasts have Multinucleate, many. The main structures making up the nucleus are the nuclear envelope, a double membrane that encloses the entire organelle and isolates its contents from the cellular cytoplasm; and the nuclear matrix, a network within the nucleus that adds mechanical support. The cell nucleus contains nearly all of the cell's genome. Nuclear DNA is often organized into multiple chromosomes – long strands of DNA dotted with various proteins, such as histones, that protect and organize the DNA. The genes within these chromosomes are Nuclear organization, structured in such a way to promote cell function. The nucleus maintains the integrity of genes and controls the activities of the ...
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