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Overlapping Generations
In population genetics overlapping generations refers to mating systems where more than one breeding generation is present at any one time. In systems where this is not the case there are non-overlapping generations (or discrete generations) in which every breeding generation lasts just one breeding season. If the adults reproduce over multiple breeding seasons the species is considered to have overlapping generations. Examples of species which have overlapping generations are many mammals, including humans, and many invertebrates in seasonal environments. Examples of species which consist of non-overlapping generations are annual plants and several insect species. Non-overlapping generations is one of the characteristics that needs to be met in the Hardy–Weinberg model for evolution to occur. This is a very restrictive and unrealistic assumption, but one that is difficult to dispose of. Overlapping versus non-overlapping generations In population genetics models, such as the ...
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Population Genetics
Population genetics is a subfield of genetics that deals with genetic differences within and among populations, and is a part of evolutionary biology. Studies in this branch of biology examine such phenomena as Adaptation (biology), adaptation, speciation, and population stratification, population structure. Population genetics was a vital ingredient in the emergence of the Modern synthesis (20th century), modern evolutionary synthesis. Its primary founders were Sewall Wright, J. B. S. Haldane and Ronald Fisher, who also laid the foundations for the related discipline of quantitative genetics. Traditionally a highly mathematical discipline, modern population genetics encompasses theoretical, laboratory, and field work. Population genetic models are used both for statistical inference from DNA sequence data and for proof/disproof of concept. What sets population genetics apart from newer, more phenotypic approaches to modelling evolution, such as evolutionary game theory and evolu ...
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Invertebrate
Invertebrates are animals that neither develop nor retain a vertebral column (commonly known as a ''spine'' or ''backbone''), which evolved from the notochord. It is a paraphyletic grouping including all animals excluding the chordata, chordate subphylum Vertebrata, i.e. vertebrates. Well-known Phylum, phyla of invertebrates include arthropods, molluscs, annelids, echinoderms, flatworms, cnidarians, and sponges. The majority of animal species are invertebrates; one estimate puts the figure at 97%. Many invertebrate taxon, taxa have a greater number and diversity of species than the entire subphylum of Vertebrata. Invertebrates vary widely in size, from 10 Micrometre, μm (0.0004 in) myxozoans to the 9–10 m (30–33 ft) colossal squid. Some so-called invertebrates, such as the Tunicata and Cephalochordata, are actually sister chordate subphyla to Vertebrata, being more closely related to vertebrates than to other invertebrates. This makes the "invertebrates" para ...
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Annual Plant
An annual plant is a plant that completes its life cycle, from germination to the production of seeds, within one growing season, and then dies. Globally, 6% of all plant species and 15% of herbaceous plants (excluding trees and shrubs) are annuals. The annual life cycle has independently emerged in over 120 different plant families throughout the entire angiosperm phylogeny. The evolutionary and ecological drivers of the annual life cycle Traditionally, there has been a prevailing assumption that annuals have evolved from perennial ancestors. However, recent research challenges this notion, revealing instances where perennials have evolved from annual ancestors. Intriguingly, models propose that transition rates from an annual to a perennial life cycle are twice as fast as the reverse transition. The life-history theory posits that annual plants are favored when adult mortality is higher than seedling (or seed) mortality, i.e., annuals will dominate environments with dis ...
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Hardy–Weinberg Principle
In population genetics, the Hardy–Weinberg principle, also known as the Hardy–Weinberg equilibrium, model, theorem, or law, states that Allele frequency, allele and genotype frequencies in a population will remain constant from generation to generation in the absence of other evolutionary influences. These influences include ''genetic drift'', ''mate choice'', ''assortative mating'', ''natural selection'', ''sexual selection'', ''mutation'', ''gene flow'', ''meiotic drive'', ''genetic hitchhiking'', ''population bottleneck'', ''founder effect,'' ''inbreeding and outbreeding depression''. In the simplest case of a single locus with two alleles denoted ''A'' and ''a'' with frequencies and , respectively, the expected genotype frequencies under random mating are for the AA homozygotes, for the aa homozygotes, and for the heterozygotes. In the absence of selection, mutation, genetic drift, or other forces, allele frequencies ''p'' and ''q'' are constant between generations ...
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Evolution
Evolution is the change in the heritable Phenotypic trait, characteristics of biological populations over successive generations. It occurs when evolutionary processes such as natural selection and genetic drift act on genetic variation, resulting in certain characteristics becoming more or less common within a population over successive generations. The process of evolution has given rise to biodiversity at every level of biological organisation. The scientific theory of evolution by natural selection was conceived independently by two British naturalists, Charles Darwin and Alfred Russel Wallace, in the mid-19th century as an explanation for why organisms are adapted to their physical and biological environments. The theory was first set out in detail in Darwin's book ''On the Origin of Species''. Evolution by natural selection is established by observable facts about living organisms: (1) more offspring are often produced than can possibly survive; (2) phenotypic variatio ...
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Altruism
Altruism is the concern for the well-being of others, independently of personal benefit or reciprocity. The word ''altruism'' was popularised (and possibly coined) by the French philosopher Auguste Comte in French, as , for an antonym of egoism. He derived it from the Italian , which in turn was derived from Latin , meaning "alterity, other people" or "somebody else". Altruism may be considered a synonym of selflessness, the opposite of self-centeredness. Altruism is an important moral value in many cultures and religions. It can Moral circle expansion, expand beyond care for humans to include other Sentience, sentient beings and future generations. Altruism, as observed in populations of organisms, is when an individual performs an action at a cost to itself (in terms of e.g. pleasure and quality of life, time, probability of survival or reproduction) that benefits, directly or indirectly, another individual, without the expectation of reciprocity or compensation for that ac ...
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Amegilla Dawsoni
''Amegilla dawsoni'', sometimes called the Dawson's burrowing bee, is a species of bee that nests by the thousands in arid claypans in Western Australia. It is a long tongued bee, of the tribe Anthophorini and genus ''Amegilla,'' the second largest genus in Anthophorini. The Dawson's burrowing bee is one of the largest Australian bee species, growing to be in length and in wingspan. With the exception of their faces, the bees are covered in brown fur, if male, or brown and white fur if female. They are similar in size and coloring to Australian carpenter bees. They are known solitary nesters. Though each female bee will build her own nest, they aggregate in large communities that give the appearance of colonies. Their nests are dug into the ground, with individual capsules created for each brood cell. Each female will only breed once in their breeding season. The males of the species are dimorphic, based on brood provisioning strategies during development. The larger mal ...
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Voltinism
Voltinism is a term used in biology to indicate the number of broods or generations of an organism in a year. The term is most often applied to insects, and is particularly in use in sericulture, where silkworm varieties vary in their voltinism. * Univoltine (monovoltine) – (adjective) referring to organisms having one brood or generation per year * Bivoltine (divoltine) – (adjective) referring to organisms having two broods or generations per year *Trivoltine – (adjective) referring to organisms having three broods or generations per year * Multivoltine (polyvoltine) – (adjective) referring to organisms having more than two broods or generations per year * Semivoltine – There are two meanings: :* (''biology'') Less than univoltine; having a brood or generation less often than once per year :* or (adjective) referring to organisms whose generation time is more than one year. Examples The speckled wood butterfly is univoltine in the northern part of its range, e.g. nort ...
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Genetic Diversity
Genetic diversity is the total number of genetic characteristics in the genetic makeup of a species. It ranges widely, from the number of species to differences within species, and can be correlated to the span of survival for a species. It is distinguished from '' genetic variability'', which describes the tendency of genetic characteristics to vary. Genetic diversity serves as a way for populations to adapt to changing environments. With more variation, it is more likely that some individuals in a population will possess variations of alleles that are suited for the environment. Those individuals are more likely to survive to produce offspring bearing that allele. The population will continue for more generations because of the success of these individuals. The academic field of population genetics includes several hypotheses and theories regarding genetic diversity. The neutral theory of evolution proposes that diversity is the result of the accumulation of neutral substitu ...
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Genetic Variance
Genetic variance is a concept outlined by the England, English biologist and statistics, statistician Ronald Fisher in his Fisher's fundamental theorem of natural selection, fundamental theorem of natural selection. In his 1930 book ''The Genetical Theory of Natural Selection'', Fisher postulates that the rate of change of fitness (biology), biological fitness can be calculated by the genetic variance of the fitness itself. Fisher tried to give a statistical formula about how the change of fitness in a population can be attributed to changes in the allele frequency. Fisher made no restrictive assumptions in his formula concerning fitness parameters, mate choices or the number of alleles and locus (genetics), loci involved. Definition Phenotypic variance, usually combines the genotype variance with the environmental variance. Genetic variance has three major components: the additive genetic variance, dominance variance, and epistatic variance. Additive genetic variance involves ...
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Genetic Drift
Genetic drift, also known as random genetic drift, allelic drift or the Wright effect, is the change in the Allele frequency, frequency of an existing gene variant (allele) in a population due to random chance. Genetic drift may cause gene variants to disappear completely and thereby reduce genetic variation. It can also cause initially rare alleles to become much more frequent and even fixed. When few copies of an allele exist, the effect of genetic drift is more notable, and when many copies exist, the effect is less notable (due to the law of large numbers). In the middle of the 20th century, vigorous debates occurred over the relative importance of natural selection versus neutral processes, including genetic drift. Ronald Fisher, who explained natural selection using Mendelian inheritance, Mendelian genetics, held the view that genetic drift plays at most a minor role in evolution, and this remained the dominant view for several decades. In 1968, population geneticist Mot ...
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Effective Population Size
The effective population size (''N''''e'') is the size of an idealised population that would experience the same rate of genetic drift as the real population. Idealised populations are those following simple one- locus models that comply with assumptions of the neutral theory of molecular evolution. The effective population size is normally smaller than the census population size ''N'', partly because chance events prevent some individuals from breeding, and partly due to background selection and genetic hitchhiking. The same real population could have a different effective population size for different properties of interest, such as genetic drift (or more precisely, the speed of coalescence) over one generation vs. over many generations. Within a species, areas of the genome that have more genes and/or less genetic recombination tend to have lower effective population sizes, because of the effects of selection at linked sites. In a population with selection at many loci an ...
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