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Opisthodactylus
''Opisthodactylus'' is an extinct genus of rheiform bird from the Early to Middle Miocene Santa Cruz and Chichinales Formations and the Late Miocene (Montehermosan) Andalhuala Formation of Argentina. Three species are described: the type species, ''O. patagonicus'', ''O. kirchneri'' and ''O. horacioperezi''. The species ''O. kirchneri'' was described in 2017 by Noriega et al.''Opisthodactylus''
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Fossilworks Fossilworks was a portal which provides query, download, and analysis tools to facilitate access to the Paleobiology Database, a large relational database assembled by hundreds of paleontologists from around the world. History Fossilworks was cr ...
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Patagorhacos
''Patagorhacos'' is an extinct genus of medium-sized phorusrhacid from the early Miocene of Patagonia. Currently only the single species ''Patagorhacos terrificus'' is known, which is represented by two highly fragmentary specimen, one belonging to the back of the skull and the other being the distal end of a leg bone. It was described together with the contemporary rheid '' Opisthodactylus horacioperezi''. History and naming Fossil remains of this phorusrhacid have been discovered in Paso Córdoba, a Natural Protection area in the Rio Negro Province of Argentina. The rocks of the area belong to the Chichinales Formation, which dates to the early Miocene (Colhuehuapian in accordance with South American land mammal age (SALMA) classification. ''Patagorhacos'' is known from only two specimens. The holotype (MPCN-PV-377) is the distal end of a right quadrate, while the single referred specimen consists of the distal end of an ulna (MPCN-PV-379). Despite the lack in overlap, the ul ...
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Santacrucian
The Santacrucian age is a period of geologic time (17.5 – 16.3 Mya (unit), Ma) within the Early Miocene epoch of the Neogene, used more specifically with South American land mammal age, SALMA classification in South America. It follows the Colhuehuapian and precedes the Friasian age. Etymology The age is named after the Santa Cruz Formation in the Magallanes Basin, Austral/Magallanes Basin of southern Patagonia, Argentina and Chile. Formations Fossils References Bibliography

;Santa Cruz Formation * * * * * * * * * * * * * * * * ;Aisol Formation * * ;Cantaure Formation * ;Castillo Formation * * * * * ;Cerro Boleadores Formation * ;Chaguaramas Formation * ;Chilcatay Formation * * * * ;Cura-Mallín Group * * * ;Gran Bajo del Gualicho Formation * ;Jimol Formation * * * * * * * * ;Mariño Formation * * * * * ;Pinturas Formation * * ;Río Yuca Formation * {{SALMA Santacrucian, Miocene South America ...
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Colhuehuapian
The Colhuehuapian age is a period of geologic time (21.0–17.5 Ma) within the Early Miocene epoch of the Neogene, used more specifically within the SALMA classification in South America. It follows the Deseadan and precedes the Santacrucian age. Etymology The age is named after the Colhué Huapí Member of the Sarmiento Formation in the Golfo San Jorge Basin, Patagonia, Argentina Argentina, officially the Argentine Republic, is a country in the southern half of South America. It covers an area of , making it the List of South American countries by area, second-largest country in South America after Brazil, the fourt .... Formations Fossils References Bibliography ;Colhué Huapí Member * * ;Abanico Formation * * * * * * ;Biblián Formation * ;Castillo Formation * * * * * ;Cerro Bandera Formation * * ;Chichinales Formation * * ;Chilcatay Formation * * * * * * * * * * ;Cura-Mallín Group * * * ;Gaiman Formati ...
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Rheidae
Rheidae is a family of flightless ratite birds which first appeared in the Paleocene. It is today represented by the sole living genus '' Rhea'', but also contains several extinct genera. Taxonomy Order Rheiformes (Forbes, 1884) Furbringer, 1888 heimorphae Bonaparte, 1849; Rheae Forbes 1884ref name="mikko"> * Family † Opisthodactylidae Ameghino 1895 ** Genus ?†'' Diogenornis'' de Alvarenga 1983 (Late Paleocene) – possibly a member of Casuariiformes The Casuariiformes is an order of large flightless birds that has four surviving members: the three species of cassowary, and the only remaining species of emu. They are divided into either a single family, Casuariidae, or occasionally two, wit ... instead. ** Genus †'' Opisthodactylus'' Ameghino 1895 (Miocene) – rheid? * Family Rheidae (Bonaparte 1849) Bonaparte, 1853 ** Genus †'' Heterorhea'' Rovereto 1914 (Pliocene) ** Genus †'' Hinasuri'' Tambussi 1995 ** Genus '' Rhea'' Brisson 1760 References {{Author ...
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Santa Cruz Formation, Argentina
The Santa Cruz Formation is a geological formation in the Magallanes Basin, Magallanes/Austral Basin in southern Patagonia in Argentina and adjacent areas of Chile. It dates to the late Early Miocene epoch, and is contemporaneous with the eponymous Santacrucian age of the SALMA (South American land mammal age) timescale. The Santa Cruz Formation is known for its abundance of vertebrate fossils, including South American native ungulates (Astrapotheria, astrapotheres, Litopterna, litopterns, Notoungulata, notoungulates), as well as rodents, xenarthrans (armadillos, sloths, anteaters), and metatherians. The formation extends from the Andes to the Atlantic coast. In its coastal section it is divided into two members, the lower, fossil rich Estancia La Costa Member, which consists predominately of tuffaceous deposits and fine grained mudrock, and the upper fossil-poor Estancia La Angelina Member, which consists of sedimentary rock, primarily mudrock, and sandstone. The environment of ...
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Chichinales Formation
The Chichinales Formation is a geological formation in Río Negro Province, Argentina which dates from the Late Oligocene to the Early Miocene, around 23 to 17.5 million years ago. It predominantly consists of pyroclastic deposits, which were deposited in a semi-arid environment. It is divided up into a number of members. The diverse fauna of the Chichinales Formation, including a variety of turtles and birds, also includes many mammals such as South American native ungulates (notoungulates and litopterns) as well as armadillos, and caviomorph rodents. It is named after the type locality and extends over a large part of the province of Río Negro. It outcrops in the valle de Río Negro, in the towns of General Roca, Río Negro and Villa Regina, bordering the southern bank of the Río Negro. Part of the most important outcrops are located within the Paso Córdova Natural Protected Area. The Chichinales Formation, broadly speaking, is made up of grayish sandstones, whitish tu ...
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Miocene
The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recent" because it has 18% fewer modern marine invertebrates than the Pliocene has. The Miocene followed the Oligocene and preceded the Pliocene. As Earth went from the Oligocene through the Miocene and into the Pliocene, the climate slowly cooled towards a series of ice ages. The Miocene boundaries are not marked by distinct global events but by regionally defined transitions from the warmer Oligocene to the cooler Pliocene Epoch. During the Early Miocene, Afro-Arabia collided with Eurasia, severing the connection between the Mediterranean and Indian Oceans, and allowing the interchange of fauna between Eurasia and Africa, including the dispersal of proboscideans and Ape, hominoids into Eurasia. During the late Miocene, the conn ...
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Fossil Taxa Described In 1891
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in amber, hair, petrified wood and DNA remnants. The totality of fossils is known as the ''fossil record''. Though the fossil record is incomplete, numerous studies have demonstrated that there is enough information available to give a good understanding of the pattern of diversification of life on Earth. In addition, the record can predict and fill gaps such as the discovery of ''Tiktaalik'' in the arctic of Canada. Paleontology includes the study of fossils: their age, method of formation, and evolutionary significance. Specimens are sometimes considered to be fossils if they are over 10,000 years old. The oldest fossils are around 3.48 billion years to 4.1 billion years old. Early edition, published online before print. The ob ...
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Neogene Argentina
The Neogene ( ,) is a geologic period and system that spans 20.45 million years from the end of the Paleogene Period million years ago ( Mya) to the beginning of the present Quaternary Period million years ago. It is the second period of the Cenozoic and the eleventh period of the Phanerozoic. The Neogene is sub-divided into two epochs, the earlier Miocene and the later Pliocene. Some geologists assert that the Neogene cannot be clearly delineated from the modern geological period, the Quaternary. The term "Neogene" was coined in 1853 by the Austrian palaeontologist Moritz Hörnes (1815–1868). The earlier term Tertiary Period was used to define the span of time now covered by Paleogene and Neogene and, despite no longer being recognized as a formal stratigraphic term, "Tertiary" still sometimes remains in informal use. During this period, mammals and birds continued to evolve into modern forms, while other groups of life remained relatively unchanged. The first humans ('' ...
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Friasian
The Friasian age is a period of geologic time (16.3–15.5 Ma) within the Early Miocene epoch of the Neogene, used more specifically within the SALMA classification of South America. It follows the Santacrucian and precedes the Colloncuran age. Etymology The age is named after the Río Frías Formation in the Aysén Basin, Patagonia, Chile Chile, officially the Republic of Chile, is a country in western South America. It is the southernmost country in the world and the closest to Antarctica, stretching along a narrow strip of land between the Andes, Andes Mountains and the Paci .... Formations Fossils References Bibliography ;Río Frías Formation * * * * * ;Castilletes Formation * * * * * * * * ;Cerdas Beds * ;Chilcatay Formation * * * * ;Cura-Mallín Group * * * ;Gran Bajo del Gualicho Formation * ;Parángula Formation * ;Pebas Formation * * ;Río Foyel Formation * * ;Río Yuca Formation * {{SALMA Miocene S ...
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Colloncuran
The Colloncuran () age is a period of geologic time (15.5–13.8 Ma) within the Middle Miocene epoch of the Neogene, used more specifically within the SALMA classification in South America. It follows the Friasian and precedes the Laventan age. Etymology The age is named after the Collón Curá Formation in the Cañadón Asfalto and Neuquén Basins of northern Patagonia, Argentina Argentina, officially the Argentine Republic, is a country in the southern half of South America. It covers an area of , making it the List of South American countries by area, second-largest country in South America after Brazil, the fourt .... Formations Fossils References Bibliography ;Collón Curá Formation * * * * * * * * * * * * * * * * * * * * * * * * * * * ;Castilletes Formation * * * * * * * * ;Cura-Mallín Group * * * ;Gran Bajo del Gualicho Formation * ;Nazareno Formation * * * * ;Pebas Formation * * ;Pisco Form ...
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