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Offacolus
''Offacolus'' is an extinct genus of euchelicerate, a group of chelicerate arthropods. Its only species, ''O. kingi'', has been found in deposits from the Silurian period ( Homerian epoch) in the Wenlock Series Lagerstätte of Herefordshire, England. The genus is named after Offa, a king from the ancient kingdom of Mercia, and ''colus'', a person who dwelled among (this time referring to) the Offa's Dyke. The species name honors Robert Joseph King, a British mineralogist who found the fossils of ''Offacolus''. Similar to ''Dibasterium'', ''Offacolus'' possess limb-like exopods (outer limb branches) on appendage II to V, a character suggest to be plesiomorphic (observable in the putative stem-chelicerate taxon Habeliida) and lost within the prosomapod clade. Classification ''Offacolus'' was originally described as an arthropod with chelicerate affinities, with detailed redescription done by Sutton ''et al.'' 2002 further suggested it to be unambiguously a chelicerate arthro ...
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Coalbrookdale Formation
Coalbrookdale Formation, earlier known as Wenlock Shale or Wenlock Shale Formation and also referred to as Herefordshire Lagerstätte in palaeontology, is a fossil-rich deposit ('' Konservat-Lagerstätte'') in Powys and Herefordshire at the England–Wales border in UK. It belongs to the Wenlock Series of the Silurian Period within the Homerian Age (about 430 million years ago). It is known for its well-preserved fossils of various invertebrate animals many of which are in their three-dimensional structures. Some of the fossils are regarded as earliest evidences and evolutionary origin of some of the major groups of modern animals. Roderick Murchison first described the geological setting of Coalbrookdale Formation by which he gave the name Silurian in 1935, referring to the Silures, a Celtic tribe of Wales. It is assigned to the Wenlock Group in 1978 based on the age of crustacean fossils found around the region. Robert J. King of the University of Leicester discovered the f ...
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Euchelicerate
The subphylum Chelicerata (from Neo-Latin, , ) constitutes one of the major subdivisions of the phylum Arthropoda. Chelicerates include the sea spiders, horseshoe crabs, and arachnids (including harvestmen, scorpions, spiders, solifuges, ticks, and mites, among many others), as well as a number of extinct lineages, such as the eurypterids (sea scorpions) and chasmataspidids. Chelicerata split from Mandibulata by the mid-Cambrian, as evidenced by stem-group chelicerates like Habeliida and '' Mollisonia'' present by this time. The surviving marine species include the four species of xiphosurans (horseshoe crabs), and possibly the 1,300 species of pycnogonids (sea spiders), if the latter are indeed chelicerates. On the other hand, there are over 77,000 well-identified species of air-breathing chelicerates, and there may be about 500,000 unidentified species. Like all arthropods, chelicerates have segmented bodies with jointed limbs, all covered in a cuticle made o ...
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Prosomapoda
Prosomapoda is a clade of Chelicerata, euchelicerates including the groups Xiphosura (horseshoe crabs) and Planaterga (a group comprising Bunodidae, bunodids, Pseudoniscidae, pseudoniscids, Chasmataspidida, chasmataspidids, Eurypterida, eurypterids and Arachnida, arachnids), as well as several basal Synziphosurina, synziphosurid genera. The clade is defined by the lack of exopods (outer branches) of prosomal appendage II-V in the adult instar, where in contrast the exopods of appendage II-V are well-developed in the non-prosomapod euchelicerates ''Offacolus'' and ''Dibasterium''. References

Prosomapoda, Middle Ordovician first appearances {{Chelicerata-stub ...
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Setapedites
''Setapedites'' is an extinct genus of Ordovician offacolid chelicerates from the Fezouata Formation of Morocco. The genus contains a single species, ''Setapedites abundantis''. Description ''Setapedites'' is roughly 6 mm long excluding the telson, with eleven body segments and a prosoma with six appendage pairs. The first appendage pairs are elongated chelicerae, with the other five being biramous appendages. The exopods of these appendages bear setae on their tips, hence the name ''Setapedites''. The opisthosoma is divided into a pre-abdomen and an abdomen. Each pre-abdomen segment has both a pair of appendages and on the tergites, leaf-shaped tergopleurae, although somite 14 and all of the abdominal somites lack appendages. The first pre-abdominal somite has a large, flap-like exopod. The abdominal somites have fused tergites and sternites forming ring-shaped structures, alongside two pairs of spines derived from the tergopleurae. Under the 11th tergite, a small, symmetri ...
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Offa
Offa ( 29 July 796 AD) was King of Mercia, a kingdom of Anglo-Saxon England, from 757 until his death in 796. The son of Thingfrith and a descendant of Eowa, Offa came to the throne after a period of civil war following the assassination of Æthelbald. Offa defeated the other claimant, Beornred. In the early years of Offa's reign, it is likely that he consolidated his control of Midland peoples such as the Hwicce and the Magonsæte. Taking advantage of instability in the kingdom of Kent to establish himself as overlord, Offa also controlled Sussex by 771, though his authority did not remain unchallenged in either territory. In the 780s he extended Mercian Supremacy over most of southern England, allying with Beorhtric of Wessex, who married Offa's daughter Eadburh, and regained complete control of the southeast. He also became the overlord of East Anglia and had King Æthelberht II of East Anglia beheaded in 794, perhaps for rebelling against him. Offa was a Christian ...
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Last Common Ancestor
A most recent common ancestor (MRCA), also known as a last common ancestor (LCA), is the most recent individual from which all organisms of a set are inferred to have descended. The most recent common ancestor of a higher taxon is generally assumed to have been a species. The term is also used in reference to the ancestry of groups of genes (haplotypes) rather than organisms. The ancestry of a set of individuals can sometimes be determined by referring to an established pedigree, although this may refer only to patrilineal or matrilineal lines for sexually-reproducing organisms with two parents, four grandparents, etc. However, in general, it is impossible to identify the exact MRCA of a large set of individuals, but an estimate of the time at which the MRCA lived can often be given. Such ''time to most recent common ancestor'' (''TMRCA'') estimates can be given based on DNA test results and established mutation rates as practiced in genetic genealogy, or by reference to a non- ...
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Sister Taxon
In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and taxon B are sister groups to each other. Taxa A and B, together with any other extant or extinct descendants of their most recent common ancestor (MRCA), form a monophyletic group, the clade AB. Clade AB and taxon C are also sister groups. Taxa A, B, and C, together with all other descendants of their MRCA form the clade ABC. The whole clade ABC is itself a subtree of a larger tree which offers yet more sister group relationships, both among the leaves and among larger, more deeply rooted clades. The tree structure shown connects through its root to the rest of the universal tree of life. In cladistic standards, taxa A, B, and C may represent specimens, species, genera, or any other taxonomic units. If A and B are at the same taxonomic ...
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Monotypic
In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispecific" or "monospecific" is sometimes preferred. In botanical nomenclature, a monotypic genus is a genus in the special case where a genus and a single species are simultaneously described. Theoretical implications Monotypic taxa present several important theoretical challenges in biological classification. One key issue is known as "Gregg's Paradox": if a single species is the only member of multiple hierarchical levels (for example, being the only species in its genus, which is the only genus in its family), then each level needs a distinct definition to maintain logical structure. Otherwise, the different taxonomic ranks become effectively identical, which creates problems for organizing biological diversity in a hierarchical syste ...
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Family (biology)
Family (, : ) is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between order and genus. A family may be divided into subfamilies, which are intermediate ranks between the ranks of family and genus. The official family names are Latin in origin; however, popular names are often used: for example, walnut trees and hickory trees belong to the family Juglandaceae, but that family is commonly referred to as the "walnut family". The delineation of what constitutes a family—or whether a described family should be acknowledged—is established and decided upon by active taxonomists. There are not strict regulations for outlining or acknowledging a family, yet in the realm of plants, these classifications often rely on both the vegetative and reproductive characteristics of plant species. Taxonomists frequently hold varying perspectives on these descriptions, leading to a lack of widespread consensus within the scientific community ...
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Homerian
In the geologic timescale, the Homerian is an age of the Wenlock Epoch of the Silurian Period of the Paleozoic Era of the Phanerozoic Eon that is comprehended between 430.5 ± 0.7 Ma and 427.4 ± 0.5 Ma (million years ago), approximately. The Homerian Age succeeds the Sheinwoodian Age and precedes the Gorstian Age. The name comes from the small village of Homer, Shropshire near Much Wenlock. The defining lower boundary of Homerian rock layers (GSSP A Global Boundary Stratotype Section and Point (GSSP), sometimes referred to as a golden spike, is an internationally agreed upon reference point on a stratigraphic section which defines the lower boundary of a stage on the geologic time scale. ...) is located within the Coalbrookdale Formation of England. References Wenlock epoch Silurian geochronology {{geochronology-stub ...
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Dibasterium
''Dibasterium'' is an extinct genus of euchelicerate, a group of chelicerate arthropods. Fossils of the single and type species, ''D. durgae'', have been discovered in the Coalbrookdale Formation of the Middle Silurian period (Homerian age) in Herefordshire, England (in the United Kingdom). The name of the genus is derived from the Latin words ''dibamos'' ("on two legs") and '' mysterium'' ("mystery"), meaning "mystery on two legs" and referring to its prosomal (of the head) limbs. The species name ''durgae'' comes from Durga, a Hindu goddess Hindu deities are the gods and goddesses in Hinduism. Deities in Hinduism are as diverse as its traditions, and a Hindu can choose to be polytheistic, pantheistic, monotheistic, monistic, even agnostic, atheistic, or humanist. Julius J. Lip ... with many arms. References Euchelicerata Fossils of England Silurian arthropods of Europe Silurian animals Wenlock first appearances Homerian extinctions Fossil taxa described in ...
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