Minjinia
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Minjinia
''Minjinia turgenensis'' is a species of placoderm from the Devonian of Mongolia. It is known from a single specimen preserving part of the skull, including remains of endochondral bone, which indicates that a mineralised endoskeleton evolved before the split between bony and cartilaginous fish, and that it was lost in the latter group. ''Minjinia'', an ancient armored fish from the Devonian period, helps scientists understand how the shoulder bones of vertebrates first developed. Researchers found evidence that its skull was connected to the shoulder in a way that suggests the shoulder bones may have evolved from parts of the gill skeleton, supporting one of the main ideas about how fish fins and shoulders first appeared.Brazeau, M. D., Castiello, M., El Fassi El Fehri, A., Hamilton, L., Ivanov, A. O., Johanson, Z., & Friedman, M. (2023, November 1). Fossil evidence for a pharyngeal origin of the vertebrate pectoral girdle. Nature News. https://www.nature.com/articles/s41586-023 ...
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Placoderm
Placoderms (from Ancient Greek πλάξ [''plax'', ''plakos''] 'Plate (animal anatomy), plate' and δέρμα [''derma''] 'skin') are vertebrate animals of the class (biology), class Placodermi, an extinct group of prehistoric fish known from Paleozoic fossils during the Silurian and the Devonian geological period, periods. While their endoskeletons are mainly cartilaginous, their head and thorax were covered by articulated armour (zoology), armoured plates (hence the name), and the rest of the body was scale (zoology), scaled or naked depending on the species. Placoderms were among the first jawed fish (their fish jaw, jaws likely Evolution, evolved from the first pair of gill arches), as well as the first vertebrates to have true tooth, teeth. They were also the first fish clade to develop pelvic fins, the second set of paired fins and the homology (biology), homologous precursor to hindlimbs in tetrapods. 380-million-year-old fossils of three other genera, ''Incisoscutum'', ''M ...
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Placoderm Genera
Placoderms (from Ancient Greek πλάξ 'plax'', ''plakos''' plate' and δέρμα 'derma'''skin') are vertebrate animals of the class Placodermi, an extinct group of prehistoric fish known from Paleozoic fossils during the Silurian and the Devonian periods. While their endoskeletons are mainly cartilaginous, their head and thorax were covered by articulated armoured plates (hence the name), and the rest of the body was scaled or naked depending on the species. Placoderms were among the first jawed fish (their jaws likely evolved from the first pair of gill arches), as well as the first vertebrates to have true teeth. They were also the first fish clade to develop pelvic fins, the second set of paired fins and the homologous precursor to hindlimbs in tetrapods. 380-million-year-old fossils of three other genera, '' Incisoscutum'', '' Materpiscis'' and '' Austroptyctodus'', represent the oldest known examples of live birth. Placoderms are thought to be paraphyletic, cons ...
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Gnathostomata
Gnathostomata (; from Ancient Greek: (') 'jaw' + (') 'mouth') are jawed vertebrates. Gnathostome diversity comprises roughly 60,000 species, which accounts for 99% of all extant vertebrates, including all living bony fishes (both ray-finned and lobe-finned, including their terrestrial tetrapod relatives) and cartilaginous fishes, as well as extinct prehistoric fish such as placoderms and acanthodians. Most gnathostomes have retained ancestral traits like true teeth, a stomach, and paired appendages ( pectoral and pelvic fins, limbs, wings, etc.). Other traits are elastin, horizontal semicircular canal of the inner ear, myelinated neurons, and an adaptive immune system which has discrete secondary lymphoid organs (spleen and thymus) and uses V(D)J recombination to create antigen recognition sites, rather than using genetic recombination in the variable lymphocyte receptor gene. It is now assumed that Gnathostomata evolved from ancestors that already possessed two pairs ...
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Osteichthyes
Osteichthyes ( ; ), also known as osteichthyans or commonly referred to as the bony fish, is a Biodiversity, diverse clade of vertebrate animals that have endoskeletons primarily composed of bone tissue. They can be contrasted with the Chondrichthyes (cartilaginous fish) and the extinct placoderms and acanthodians, which have endoskeletons primarily composed of cartilage. The vast majority of extant taxon, extant fish are members of Osteichthyes, being an extremely diverse and abundant group consisting of 45 order (biology), orders, over 435 family (biology), families and 28,000 species. The group is divided into two main clades, the ray-finned fish (Actinopterygii, which makes up the vast majority of extant fish) and the lobe-finned fish (Sarcopterygii, which gave rise to all land vertebrates, i.e. tetrapods). The oldest known fossils of bony fish are about 425 million years old from the late Silurian, which are also transitional fossils showing a dentition, tooth pattern th ...
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Chondrichthyes
Chondrichthyes (; ) is a class of jawed fish that contains the cartilaginous fish or chondrichthyans, which all have skeletons primarily composed of cartilage. They can be contrasted with the Osteichthyes or ''bony fish'', which have skeletons primarily composed of bone tissue. Chondrichthyes are aquatic vertebrates with paired fins, paired nares, placoid scales, conus arteriosus in the heart, and a lack of opercula and swim bladders. Within the infraphylum Gnathostomata, cartilaginous fishes are distinct from all other jawed vertebrates. The class is divided into two subclasses: Elasmobranchii (sharks, rays, skates and sawfish) and Holocephali ( chimaeras, sometimes called ghost sharks, which are sometimes separated into their own class). Extant chondrichthyans range in size from the finless sleeper ray to the over whale shark. Anatomy Skeleton The skeleton is cartilaginous. The notochord is gradually replaced by a vertebral column during development, e ...
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Eugnathostomata
Gnathostomata (; from Ancient Greek: (') 'jaw' + (') 'mouth') are jawed vertebrates. Gnathostome diversity comprises roughly 60,000 species, which accounts for 99% of all extant vertebrates, including all living bony fishes (both ray-finned and lobe-finned, including their terrestrial tetrapod relatives) and cartilaginous fishes, as well as extinct prehistoric fish such as placoderms and acanthodians. Most gnathostomes have retained ancestral traits like true teeth, a stomach, and paired appendages (pectoral and pelvic fins, limbs, wings, etc.). Other traits are elastin, horizontal semicircular canal of the inner ear, myelinated neurons, and an adaptive immune system which has discrete secondary lymphoid organs (spleen and thymus) and uses V(D)J recombination to create antigen recognition sites, rather than using genetic recombination in the variable lymphocyte receptor gene. It is now assumed that Gnathostomata evolved from ancestors that already possessed two pairs of ...
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Arthrodira
Arthrodira (Greek for "jointed neck") is an order of extinct armored, jawed fishes of the class Placodermi that flourished in the Devonian period before their sudden extinction, surviving for about 50 million years and penetrating most marine ecological niches. Arthrodires were the largest and most diverse of all groups of placoderms. Description Arthrodire placoderms are notable for the movable joint between armor surrounding their heads and bodies. Like all placoderms, they lacked distinct teeth; instead, they used the sharpened edges of a bony plate on their jawbone as a biting surface. The eye sockets are covered by a bony ring, which supports the eye, a feature shared by birds and some ichthyosaurs. Early arthrodires, such as the genus '' Arctolepis'', were well-armoured fishes with flattened bodies. The largest member of this group, ''Dunkleosteus'', was a true superpredator of the latest Devonian period, reaching as much as 6 m in length. In contrast, the long-nosed '' Ro ...
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Ptyctodontida
The ptyctodontids ("folded-teeth") are placoderms of the order Ptyctodontida, containing the family Ptyctodontidae. With their big heads, big eyes, reduced armor and long bodies, the ptyctodontids bore a superficial resemblance to modern day chimaeras (Holocephali). Their armor was reduced to a pattern of small plates around the head and neck. Like the extinct and related acanthothoracids, and the living and unrelated holocephalians, most of the ptyctodontids are thought to have lived near the sea bottom and preyed on shellfish. On account of their radically reduced armor, some paleontologists have suggested that the Ptyctodontida were not actually placoderms, but actual holocephalians, some primitive group of elasmobranch fish, or even were the ancestors of the holocephalians, including the chimaeras. Thorough anatomical examinations of whole fossil specimens reveal that the profound similarities between these two groups are actually very superficial. The major differences ...
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Petalichthyida
Petalichthyida is an Extinction (biology), extinct Order (biology), order of small, flattened placoderm fish. They are typified by their splayed pectoral fins, exaggerated lateral spines, flattened bodies, and numerous tubercles that decorated all of the plates and scales of their armor. They reached a peak in diversity during the Early Devonian and were found throughout the world, particularly in Europe (especially in Germany), North America, Asia, South America, and Australia. The petalichthids ''Lunaspis'' and ''Wijdeaspis'' are among the best known. The earliest and most primitive known petalichthyid is ''Diandongpetalichthys'', which is from earliest Devonian-aged strata of Yunnan. The presence of ''Diandongpetalichthys'', along with other primitive petalichthyids including ''Neopetalichthys'' and ''Quasipetalichthys'', and more advanced petalichthyids, suggest that the order may have arisen in China, possibly during the late Silurian. Because they had compressed body fo ...
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Rhenanida
Rhenanida ("Rhine (fish)") is an order of scaly placoderms. Unlike most other placoderms, the rhenanids' armor was made up of a mosaic of unfused scales and tubercles. The patterns and components of this "mosaic" correspond to the plates of armor in other, more advanced placoderms, suggesting that the ancestral placoderm had armor made of unfused components, as well. All rhenanids were flattened, ray-like, bottom-dwelling predators that lived in marine environments. Evolution The rhenanids were once presumed to be the most primitive, or at least the closest to the ancestral placoderm, as their armor was made up of a mosaic of tubercles, as opposed to the solidified plates of "advanced" placoderms, such as antiarchs and arthrodires. However, comparing the skull anatomies of ''Jagorina pandora'' with those of antiarchs, the rhenanids are considered to be the sister group of the antiarchs (together with their respective Acanthothoracid relatives). Presence in the fossil record ...
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Antiarchi
Antiarchi ("opposite anus") is an order of heavily armored placoderms. The antiarchs form the second-most successful group of placoderms after the arthrodires in terms of numbers of species and range of environments. The order's name was coined by Edward Drinker Cope, who, when examining some fossils that he thought were armored tunicates related to '' Chelyosoma'', mistakenly thought that the orbital fenestra (i.e., the hole in the headshield for the eyes, nose and pineal foramen) was the opening for the mouth, or oral siphon, and that the opening for the anal siphon was on the other side of the body, as opposed to having both oral and anal siphons together at one end. The front portions of their bodies were heavily armored, to the point of literally resembling a box with eyes, with the sometimes scaled, sometimes naked rear portions often becoming sinuous, particularly with later forms. The pair of pectoral fins were modified into a pair of caliper-like, or arthropod-like l ...
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Osteostraci
The class Osteostraci (meaning "bony shells") is an extinct taxon of bony-armored jawless fish, termed " ostracoderms", that lived in what is now North America, Europe and Russia from the Middle Silurian to Late Devonian. Anatomically speaking, the osteostracans, especially the Devonian species, were among the most advanced of all known agnathans. This is due to the development of paired fins, and their complicated cranial anatomy. The osteostracans were more similar to lampreys than to jawed vertebrates in possessing two pairs of semicircular canals in the inner ear, as opposed to the three pairs found in the inner ears of jawed vertebrates. They are thought to be the sister-group to pituriaspids, and together, these two taxa of jawless vertebrates are the sister-group of gnathostomes. Several synapomorphies support this hypothesis, such as the presence of: sclerotic ossicles, paired pectoral fins, a dermal skeleton with three layers (a basal layer of isopedin, a middle layer ...
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