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Machairodonts
Machairodontinae (from Ancient Greek μάχαιρα ''machaira,'' a type of Ancient Greek sword and ὀδόντος ''odontos'' meaning tooth) is an extinct subfamily of carnivoran mammals of the cat family Felidae, representing the earliest diverging major branch of the family. Machairodonts varied in size from comparable to lynxes to exceeding that of lions. The Machairodontinae contain many of the extinct predators commonly known as "saber-toothed cats", including those with greatly elongated upper maxillary canines, such as the famed genus ''Smilodon'' and ''Megantereon,'' though the degree of elongation was variable, and in some machairodontines like ''Dinofelis'' the length of the upper canines was much more modest. Sometimes, other carnivorous mammals with elongated teeth are also called saber-toothed cats, although they do not belong to the felids. Besides the machairodonts, other saber-toothed predators also arose in the nimravids, barbourofelids, machaeroidines, ...
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Saber-toothed Predator
A saber-tooth (alternatively spelled sabre-tooth) is any member of various extinct groups of predatory therapsids, predominantly carnivoran mammals, that are characterized by long, curved saber-shaped canine teeth which protruded from the mouth when closed. Among the earliest animals that can be described as "sabertooths" are the gorgonopsids, a group of non-mammalian therapsids that lived during the Middle-Late Permian, around 270-252 million years ago. Saber-toothed mammals have been found almost worldwide from the Eocene epoch to the end of the Pleistocene epoch (42 million years ago – 11,000 years ago). One of the best-known genera is the machairodont or "saber-toothed cat" ''Smilodon'', the species of which, especially ''S. fatalis'', are popularly referred to as "saber-toothed tigers", although they are not closely related to tigers (''Panthera''). Despite some similarities, not all saber-tooths are closely related to saber-toothed cats or felids in-general. Instead, ...
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Smilodon
''Smilodon'' is an extinct genus of Felidae, felids. It is one of the best known saber-toothed predators and prehistoric mammals. Although commonly known as the saber-toothed tiger, it was not closely related to the tiger or other modern cats, belonging to the extinct subfamily Machairodontinae, with an estimated date of Speciation, divergence from the ancestor of living cats around 20 million years ago. ''Smilodon'' was one of the last surviving machairodonts alongside ''Homotherium''. ''Smilodon'' lived in the Americas during the Pleistocene to early Holocene epoch (2.5 Year#mya, mya – at latest 8,200 years ago). The genus was named in 1842 based on fossils from Brazil; the generic name means or combined with . Three species are recognized today: ''S. gracilis'', ''S. fatalis'', and ''S. populator''. The two latter species were probably descended from ''S. gracilis'', which itself probably evolved from ''Megantereon''. The hundreds of specimens obtained from the La Brea T ...
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Machairodus
''Machairodus'' (from , 'knife' and 'tooth') is a genus of large Machairodontinae, machairodont or ''saber-toothed cat'' that lived in Africa and Eurasia during the Middle Miocene, Middle to Late Miocene, from 12.5 million to 8.7 million years ago. It is the animal from which the subfamily Machairodontinae gets its name. The type species of the genus, ''M. aphanistus'', was comparable to tigers in size making it an apex predator of the ecosystems it inhabited. It is currently usually placed as one of the most basal members of the tribe Homotherini, and the ancestor of later members of the tribe. History of research and taxonomy ''Machairodus'' was first named in 1832, by German Naturalist Johann Jakob Kaup. Though its remains had been known since 1824, it was believed by Georges Cuvier that the fossils had come from a species of bear, which he called ''Ursus cultridens'' (known today as ''Megantereon'') based on composite sample of teeth from different countries, species and g ...
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Smilodontini
Smilodontini is an extinct tribe within the Machairodontinae or "saber-toothed cat" subfamily of the Felidae. The tribe is also known as the "dirk-toothed cats". They were found in South America, North America, Europe, Asia, and Africa during the Middle Miocene to early Holocene, from 10.3 mya—8,200 years ago. As the name suggests, the famous genus of ''Smilodon'' is part of this group, but there are also three other distinct genera in Smilodontini. The earliest known genus of the tribe is '' Promegantereon'', once thought to be a species of '' Paramachairodus''. The tribe only became extinct 10,000 years ago, with the demise of ''Smilodon''. All of these are saber-toothed cats, meaning that they have long, narrow upper canines and stocky, well muscled proportions. Description The various members of Smilodontini are generally distinguished by their long, narrow upper canines. However, the stockiness often seen as one of the tribe's signature features varied between the differ ...
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Nimravidae
Nimravidae is an extinct family (biology), family of carnivorans, sometimes known as false saber-toothed cats, whose fossils are found in North America and Eurasia. Not considered to belong to the true cats (family Felidae), the nimravids are generally considered closely related and classified as a distinct family in the suborder Feliformia. Fossils have been dated from the Middle Eocene through the Late Oligocene epochs (Bartonian through Chattian stages, 41.03–25.9 million years ago), spanning about . The barbourofelids, which were formerly classified as a subfamily of the Nimravidae, were reassigned to their own distinct family Barbourofelidae in 2004. However in the 2020s, some studies suggest the barbourofelids are a branch of the nimravids, suggesting that this debate might not be settled yet. If accepted, the family would have persisted until the Late Miocene, and would extend their spatial distribution to Africa. Taxonomy The family Nimravidae was named by American pale ...
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Dinofelis
''Dinofelis'' is an extinct genus of machairodontine (sabre-toothed cat), usually classified in the tribe Metailurini. It was widespread in Europe, Asia, Africa and North America from 5 million to about 1.2 million years ago (early Pliocene to early Pleistocene). Fossils very similar to ''Dinofelis'' from Lothagam range back to around 8 million years ago, in the Late Miocene. Discovery and naming The genus ''Dinofelis'' was originally named by Otto Zdansky in 1924 for the species ''Dinofelis abeli''. Further fossil species were named, including ''Felis diastemata'' and ''Meganthereon barlowi'', which were later transferred to the genus ''Therailurus'', which was in turn later considered a junior synonym of ''Dinofelis''. A comprehensive review of the genus was published in 2001 by paleontologists Lars Werdelin and Margaret E. Lewis, including mentions of the then-unnamed Langebaanweg and Lothagam species, as well as naming a new species ''Dinofelis aronoki''; the species ep ...
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Homotherium
''Homotherium'' is an extinct genus of Homotherini, scimitar-toothed cat belonging to the extinct subfamily Machairodontinae that inhabited North America, Eurasia, and Africa, as well as possibly South America during the Pliocene and Pleistocene epochs from around 4 million to 12,000 years ago. It was one of the last surviving members of the subfamily alongside the more famous sabertooth ''Smilodon'', to which it was not particularly closely related. It was a large cat, comparable in size to a lion, functioning as an apex predator in the ecosystems it inhabited. It had an elongate neck and relatively elongate legs, a relatively short back and a very short tail, with the mummy of a ''H. latidens'' cub of Late Pleistocene age found in Siberia having a plain dark brown coat colour. In comparison to ''Smilodon'', the canines of ''Homotherium'' were shorter, though still longer than those of living cats, and it is suggested to have had a different ecology from ''Smilodon'' as a pursuit ...
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Metailurini
Metailurini is an extinct taxonomic tribe of large saber-toothed cats that lived in Africa, Asia, Europe, and North America from the Miocene to the Pleistocene. The best known Metalurini genera are '' Dinofelis'' and '' Metailurus''. Metailurini had canines longer than ''Neofelis'', but smaller than true saber toothed cats. The teeth were also more conical than flat, so called "scimitar-toothed", having broad and mildly elongated upper canines. Like most extinct cats, the majority of species in Metailurini are known primarily from fragments. However, the systematic position and taxonomy of these creatures is now accepted as being true members of Felidae and descended from ''Proailurus'' and ''Pseudaelurus''. Within Felidae, they had been traditionally considered to belong in Machairodontinae, albeit some have in the past proposed a relationship to Pantherinae,Lars W. van den Hoek Ostende, Michael Morlo, Doris Nagel: ''Fossils explained 52 Majestic killers: the sabre-toothed cats. ...
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Hyaenodonta
Hyaenodonta (" hyena teeth") is an extinct order of hypercarnivorous placental mammals of clade Pan-Carnivora from mirorder Ferae. Hyaenodonts were important mammalian predators that arose during the early Paleocene in Europe and persisted well into the late Miocene. Characteristics Hyaenodonts are characterized by long, often disproportionately large skulls, slender jaws, and slim bodies. They generally ranged in size from 30 to 140 cm at the shoulder. While '' Simbakubwa kutokaafrika'' may have been up to (surpassing the modern polar bear in size), this estimate is suspect due to being based on skull-body size ratios derived from felids, which have much smaller skulls for their body size. Other large hyaenodonts include two close and later-surviving relatives of ''Simbakubwa'', '' Hyainailouros'' and '' Megistotherium'' (the latter likely being the largest in the group), and the much earlier-living '' Hyaenodon gigas'' (the largest species from genus '' Hyaenodo ...
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Pantherinae
The Pantherinae is a subfamily of the Felidae; it was named and first described by Reginald Innes Pocock in 1917 as only including the ''Panthera'' species, but later also came to include the clouded leopards (genus ''Neofelis''). The Pantherinae genetically diverged from a common ancestor between and . Characteristics Pantherinae species are characterised by an imperfectly ossified hyoid bone with elastic tendons that enable their larynx to be mobile. They have a flat rhinarium that only barely reaches the dorsal side of the nose. The area between the nostrils is narrow, and not extended sidewards as in the Felinae. The ''Panthera'' species have a single, rounded, vocal fold with a thick mucosal lining, a large vocalis muscle, and a large cricothyroid muscle with long and narrow membranes. A vocal fold that is longer than enables all but the snow leopard among them to roar, as it has shorter vocal folds of that provide a lower resistance to airflow; this distinction w ...
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Pleistocene
The Pleistocene ( ; referred to colloquially as the ''ice age, Ice Age'') is the geological epoch (geology), epoch that lasted from to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was finally confirmed in 2009 by the International Union of Geological Sciences, the cutoff of the Pleistocene and the preceding Pliocene was regarded as being 1.806 million years Before Present (BP). Publications from earlier years may use either definition of the period. The end of the Pleistocene corresponds with the end of the last glacial period and also with the end of the Paleolithic age used in archaeology. The name is a combination of Ancient Greek () 'most' and (; Latinized as ) 'new'. The aridification and cooling trends of the preceding Neogene were continued in the Pleistocene. The climate was strongly variable depending on the glacial cycle, oscillating between cold Glacial period, glacial periods and warmer Interglacial, int ...
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Zanclean
The Zanclean is the lowest stage or earliest age on the geologic time scale of the Pliocene. It spans the time between 5.332 ± 0.005 Ma (million years ago) and 3.6 ± 0.005 Ma. It is preceded by the Messinian Age of the Miocene Epoch, and followed by the Piacenzian Age. The Zanclean can be correlated with regionally used stages, such as the Opoitian of New Zealand, and the Tabianian or Dacian of Central Europe. It also corresponds to the late Hemphillian to mid-Blancan North American Land Mammal Ages. In California, the Zanclean roughly corresponds to the middle part of the Delmontian stage. Definition The Zanclean Stage was introduced by Giuseppe Seguenza in 1868. It is named after '' Zancle'', the pre- Roman name for the Italian city of Messina on Sicily. The base of the Zanclean (and the Pliocene Series) lies with the top of magnetic chronozone Cr3 (about 100,000 years before the Thvera normal subchronozone C3n.4n). The base is also close to the extinction level of th ...
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