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Lutungutali
''Lutungutali'' (meaning "high hip" in the Bemba language) is an extinct genus of silesaurid dinosauriform from the Middle Triassic of Zambia. The single type species of the genus is ''Lutungutali sitwensis''. ''Lutungutali'' was named in 2013 and described from a fossil specimen, holotype NHCC LB32, including hip bones and tail vertebrae. The specimen was collected in 2009 from the upper Ntawere Formation, which dates to the Anisian stage of the Middle Triassic. ''Lutungutali'' is the first known silesaurid from Zambia and, along with the Tanzanian silesaurid '' Asilisaurus'' and dinosauriform ''Nyasasaurus'', the oldest bird-line archosaur known from body fossils (i.e. parts of the skeleton). Description ''Lutungutali'' is known from parts of its pelvis and from four caudal vertebrae that make up the base of the tail. These bones represent one individual, although bones that may belong to ''Lutungutali'' representing eight other individuals were collected from the same fossi ...
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Ntawere Formation
The Ntawere Formation is a Middle Triassic (Anisian) geological formation in Zambia, preserving fossils of synapsids, archosaurs, and temnospondyls. Geology Several different facies are present in the Ntawere Formation, reconstructing a floodplain environment. The coarsest facies are trough cross-bedded conglomeratic sandstone full of mineral concretions. These coarse deposits formed in ancient channels such as riverbeds. Another type of facies involves thick beds of mudstone interbedding with thinner layers of fine-grained sandstone, indicating alternating low- and high-energy water flow within the channels. Graded sandstone to mudstone overbank deposits (complete with ripple marks) occur near channel deposits. Extensive successions of laminated or massive mudstone are common, often containing slickensides, calcareous nodules or layers ( marls), and/or hematite nodules. These types of thick mud/marl layers likely formed in more quiet parts of the floodplain isolated from tur ...
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Silesaurid
Silesauridae is an extinct family of Triassic dinosauriforms. It is most commonly considered to be a clade of non-dinosaur dinosauriforms, and the sister group of dinosaurs. Some studies have instead suggested that most or all silesaurids comprised an early diverging clade or a paraphyletic grade within ornithischian dinosaurs. Silesaurids have a consistent general body plan, with a fairly long neck and legs and possibly quadrupedal habits, but most silesaurids are heavily fragmentary nonetheless. Furthermore, they occupied a variety of ecological niches, with early silesaurids (such as '' Lewisuchus'') being carnivorous and later taxa (such as ''Kwanasaurus'') having adaptations for specialized herbivory. As indicated by the contents of referred coprolites, ''Silesaurus'' may have been insectivorous, feeding selectively on small beetles and other arthropods. Classification Silesauridae is typically considered the sister group to Dinosauria. The group was named in 2010 by Max ...
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Middle Triassic
In the geologic timescale, the Middle Triassic is the second of three epoch (geology), epochs of the Triassic period (geology), period or the middle of three series (stratigraphy), series in which the Triassic system (stratigraphy), system is divided in chronostratigraphy. The Middle Triassic spans the time between annum, Ma and Ma (million years ago). It is preceded by the Early Triassic Epoch and followed by the Late Triassic Epoch. The Middle Triassic is divided into the Anisian and Ladinian age (geology), ages or stage (stratigraphy), stages. Formerly the middle series in the Triassic was also known as Muschelkalk. This name is now only used for a specific unit of stratum, rock strata with approximately Middle Triassic age, found in western Europe. Middle Triassic life Following the Permian–Triassic extinction event, the most devastating of all mass-extinctions, life recovered slowly. In the Middle Triassic, many groups of organisms reached higher diversity again, s ...
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Fossa (anatomy)
In anatomy, a fossa (; : fossae ( or ); ) is a depression or hollow, usually in a bone, such as the hypophyseal fossa (the depression in the sphenoid bone).Venieratos D, Anagnostopoulou S, Garidou A., A new morphometric method for the sella turcica and the hypophyseal fossa and its clinical relevance.;Folia Morphol (Warsz). 2005 Nov;64(4):240-7. Some examples include: In the skull: * Cranial fossa ** Anterior cranial fossa ** Middle cranial fossa *** Interpeduncular fossa ** Posterior cranial fossa * Hypophyseal fossa * Temporal bone fossa ** Mandibular fossa ** Jugular fossa * Infratemporal fossa * Pterygopalatine fossa * Pterygoid fossa * Lacrimal fossa ** Fossa for lacrimal gland ** Fossa for lacrimal sac * Scaphoid fossa * Condyloid fossa * Rhomboid fossa In the mandible: * Retromolar fossa In the torso: * Fossa ovalis (heart) * Infraclavicular fossa * Pyriform fossa * Substernal fossa * Iliac fossa * Ovarian fossa * Paravesical fossa * Coccygeal f ...
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Basal (phylogenetics)
In phylogenetics, basal is the direction of the ''base'' (or root) of a phylogenetic tree#Rooted tree, rooted phylogenetic tree or cladogram. The term may be more strictly applied only to nodes adjacent to the root, or more loosely applied to nodes regarded as being close to the root. Note that extant taxa that lie on branches connecting directly to the root are not more closely related to the root than any other extant taxa. While there must always be two or more equally "basal" clades sprouting from the root of every cladogram, those clades may differ widely in taxonomic rank, Phylogenetic diversity, species diversity, or both. If ''C'' is a basal clade within ''D'' that has the lowest rank of all basal clades within ''D'', ''C'' may be described as ''the'' basal taxon of that rank within ''D''. The concept of a 'key innovation' implies some degree of correlation between evolutionary innovation and cladogenesis, diversification. However, such a correlation does not make a given ca ...
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Manda Beds
The Manda Formation (also known as the Manda Beds) is a Middle Triassic (Anisian?) or possibly Late Triassic (Carnian?) geologic formation in Tanzania. It preserves fossils of many terrestrial vertebrates from the Triassic, including some of the earliest dinosauromorph archosaurs. The formation is often considered to be Anisian in age according to general tetrapod biochronology hypotheses and correlations to the ''Cynognathus'' Assemblage Zone of South Africa. However, some recent studies cast doubt to this age, suggesting that parts deposits may actually be younger (Carnian) in age. History of study One of the first to study rocks of the Manda Formation was British geologist G. M. Stockley. In 1932, Stockley explored the geology of the Ruhuhu Basin in Tanzania. He called a series of layers dating from the Late Carboniferous to the Middle Triassic the Songea Series and divided it into eight units labelled K1-K8. Stockley was also the first to describe fossils from these rocks, ...
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Late Triassic
The Late Triassic is the third and final epoch (geology), epoch of the Triassic geologic time scale, Period in the geologic time scale, spanning the time between annum, Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch and followed by the Early Jurassic Epoch. The corresponding series (stratigraphy), series of rock beds is known as the Upper Triassic. The Late Triassic is divided into the Carnian, Norian and Rhaetian Geologic time scale, ages. Many of the first dinosaurs evolved during the Late Triassic, including ''Plateosaurus'', ''Coelophysis'', ''Herrerasaurus'', and ''Eoraptor''. The Triassic–Jurassic extinction event began during this epoch and is one of the five major mass extinction events of the Earth. Etymology The Triassic was named in 1834 by Friedrich August von Namoh, Friedrich von Alberti, after a succession of three distinct rock layers (Greek meaning 'triad') that are widespread in southern Germany: the lower Buntsandstein (colourful ...
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Synapomorphy
In phylogenetics, an apomorphy (or derived trait) is a novel Phenotypic trait, character or character state that has evolution, evolved from its ancestral form (or Plesiomorphy and symplesiomorphy, plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxon, taxa and is therefore Hypothesis#Scientific hypothesis, hypothesized to have evolved in their most recent common ancestor. ) In cladistics, synapomorphy implies Homology (biology), homology. Examples of apomorphy are the presence of Terrestrial locomotion#Posture, erect gait, fur, Evolution of mammalian auditory ossicles, the evolution of three middle ear bones, and mammary glands in mammals but not in other vertebrate animals such as amphibians or reptiles, which have retained their ancestral traits of a Terrestrial locomotion#Posture, sprawling gait and lack of fur. Thus, these derived traits are also synapomorphies of mammals in general as they are not shared by other vertebrate animals. Etymology The word ...
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Phylogenetic
In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical data and observed heritable traits of DNA sequences, protein amino acid sequences, and morphology. The results are a phylogenetic tree—a diagram depicting the hypothetical relationships among the organisms, reflecting their inferred evolutionary history. The tips of a phylogenetic tree represent the observed entities, which can be living taxa or fossils. A phylogenetic diagram can be rooted or unrooted. A rooted tree diagram indicates the hypothetical common ancestor of the taxa represented on the tree. An unrooted tree diagram (a network) makes no assumption about directionality of character state transformation, and does not show the origin or "root" of the taxa in question. In addition to their use for inferring phylogenetic pa ...
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Family (biology)
Family (, : ) is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between order and genus. A family may be divided into subfamilies, which are intermediate ranks between the ranks of family and genus. The official family names are Latin in origin; however, popular names are often used: for example, walnut trees and hickory trees belong to the family Juglandaceae, but that family is commonly referred to as the "walnut family". The delineation of what constitutes a family—or whether a described family should be acknowledged—is established and decided upon by active taxonomists. There are not strict regulations for outlining or acknowledging a family, yet in the realm of plants, these classifications often rely on both the vegetative and reproductive characteristics of plant species. Taxonomists frequently hold varying perspectives on these descriptions, leading to a lack of widespread consensus within the scientific community ...
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Autapomorphy
In phylogenetics, an autapomorphy is a distinctive feature, known as a Synapomorphy, derived trait, that is unique to a given taxon. That is, it is found only in one taxon, but not found in any others or Outgroup (cladistics), outgroup taxa, not even those most closely related to the focal taxon (which may be a species, family (biology), family or in general any clade). It can therefore be considered as an apomorphy in relation to a single taxon. The word ''autapomorphy'', introduced in 1950 by German entomology, entomologist Willi Hennig, is derived from the Greek language, Greek words αὐτός, ''autos'' "self"; ἀπό, ''apo'' "away from"; and μορφή, ''morphḗ'' = "shape". Discussion Because autapomorphies are only present in a single taxon, they do not convey information about relationship. Therefore, autapomorphies are not useful to infer phylogenetic relationships. However, autapomorphy, like synapomorphy and plesiomorphy is a relative concept depending on the ta ...
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