Loricicaris
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Loricicaris
''Loricicaris'' is a genus of extinct hymenocarine arthropod with a single species, ''Loricicaris spinocaudatus'' from the Burgess Shale, British Columbia, Canada described in 2013. Description The longest individuals reach around in length. The front of the body was covered in a convex bivalved carapace which enveloped around 65% of the total body length. The head bore a pair of robust antennae, which was made up of at least 10 segments/podomeres, with the antennae segments bearing setae (hair-like structures). The head possibly bore a pair of stalked eyes. The head morphology is poorly known, but may have been made up of at minimum three segments with an anterior sclerite, whether a hypostome was present is unclear. The head also bore a pair of grasping appendages similar in morphology to those of ''Branchiocaris.'' Running through the body was a straight gut tube. The body was made up of somites/segments, of which 14-16 pairs were present under the carapace and 32 pairs ran ...
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Hymenocarina
Hymenocarina is an Order (biology), order of extinct marine arthropods known from the Cambrian. They possess bivalved carapaces, typically with exposed posteriors. Members of the group are morphologically diverse and had a variety of ecologies, including as filter feeders and as predators. Recent research has generally considered them to be stem or crown group members of Mandibulata, due to the presence of Mandible (arthropod mouthpart), mandibles in well-preserved species. Taxonomy Hymenocarines are characterized by the combination of the following characters: bivalved, convex carapace covering Cephalothorax, cephalothoracic (combined head and thorax) region; cephalothorax bearing multisegmented antennae (though as an exception antennae are absent in ''Odaraia'') and rounded Mandible (arthropod mouthpart), mandibles and likely Maxilla (arthropod mouthpart), maxillae, post maxillae limbs with spiny, subdivided basis and Arthropod leg, endopods (lower, leg-like parts) with well-d ...
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Protocarididae
Hymenocarina is an order of extinct marine arthropods known from the Cambrian. They possess bivalved carapaces, typically with exposed posteriors. Members of the group are morphologically diverse and had a variety of ecologies, including as filter feeders and as predators. Recent research has generally considered them to be stem or crown group members of Mandibulata, due to the presence of mandibles in well-preserved species. Taxonomy Hymenocarines are characterized by the combination of the following characters: bivalved, convex carapace covering cephalothoracic (combined head and thorax) region; cephalothorax bearing multisegmented antennae (though as an exception antennae are absent in ''Odaraia'') and rounded mandibles and likely maxillae, post maxillae limbs with spiny, subdivided basis and endopods (lower, leg-like parts) with well-developed terminal claws; absence of appendages between antennae and mandibles; median sclerite and lobate protrusions located between compou ...
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Cambrian
The Cambrian ( ) is the first geological period of the Paleozoic Era, and the Phanerozoic Eon. The Cambrian lasted 51.95 million years from the end of the preceding Ediacaran period 538.8 Ma (million years ago) to the beginning of the Ordovician Period 486.85 Ma. Most of the continents lay in the southern hemisphere surrounded by the vast Panthalassa Ocean. The assembly of Gondwana during the Ediacaran and early Cambrian led to the development of new convergent plate boundaries and continental-margin arc magmatism along its margins that helped drive up global temperatures. Laurentia lay across the equator, separated from Gondwana by the opening Iapetus Ocean. The Cambrian marked a profound change in life on Earth; prior to the Period, the majority of living organisms were small, unicellular and poorly preserved. Complex, multicellular organisms gradually became more common during the Ediacaran, but it was not until the Cambrian that fossil diversity seems to rapidly ...
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Branchiocaris
''Branchiocaris'' is an extinct genus of Cambrian bivalved arthropod. The type and best known species, ''Branchiocaris pretiosa,'' was described from the Burgess Shale of British Columbia, Canada, in 1929, originally placed in '' Protocaris'', and was placed into its own distinct genus by Briggs in 1976. Several other possible species have been described from Cambrian deposits in China, and it is also possibly known from Cambrian deposits in Utah. ''Branchiocaris pretiosa'' is around in length, with a highly segmented trunk, consisting of at least 44 ring-like segments, terminating in a forked tail telson. At the front of the animal is a pair of short segmented tapered antennules with at least 20 segments, as well as a pair of claw appendages. It was likely an active swimmer, and used the claw appendages to bring food to the mouth. The discovery of '' Tokummia'' from the Burgess Shale, believed to be a close relative of ''Branchiocaris'', has shed light on the evolutionary plac ...
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Protocaris
''Protocaris marshi'' is an extinct species of bivalved arthropod known from a single specimen collected from the Cambrian Series 2 aged Parker Formation from the Parker Quarry of northwestern Vermont, United States The specimen is preserved in top-down view and has a bivalved carapace, a segmented trunk and a forked tail. Its precise taxonomic position is uncertain, due to the limited nature of known remains, but it is suggested to be a member of Hymenocarina belonging to the family Protocarididae, which also includes '' Tokummia'' and ''Branchiocaris ''Branchiocaris'' is an extinct genus of Cambrian bivalved arthropod. The type and best known species, ''Branchiocaris pretiosa,'' was described from the Burgess Shale of British Columbia, Canada, in 1929, originally placed in '' Protocaris'', a ....'' References Cambrian arthropods Hymenocarina {{paleo-crustacean-stub ...
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Tokummia
''Tokummia'' is a genus of fossil hymenocarine arthropod, known only by one species, ''Tokummia'' ''katalepsis'', from the middle Cambrian (508 million years old) Burgess Shale as found in a quarry in Marble Canyon in Canada. Etymology The genus name ''Tokummia'' named after Tokumm Creek which runs through the Marble Canyon where it was found. The species name ''katalepsis'' is a Greek word for "seizing", "gasping" or "holding". Morphology ''Tokummia'' has a cylindrical body, with the anterior half covered by a long bivalved carapace. At the front of the animal, there are a pair of antennae, possible eyes, mouthparts (mandibles, maxillule and maxilla) and prominent pincer-like maxillipeds. These shows the oldest record of arthropod pincers. Posterior to the maxillipeds are 50 leg-bearing trunk segments. Each of its biramous leg has 5-segmented basipods, followed by an exopod (flap-like outer branch) and 7-segmented endopod (leg-like inner branch). The anterior 10 leg pair ...
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Telson
The telson () is the hindmost division of the body of an arthropod. Depending on the definition, the telson is either considered to be the final segment (biology), segment of the arthropod body, or an additional division that is not a true segment on account of not arising in the embryo from teloblast areas as other segments. It never carries any appendages, but a forked "tail" called the caudal furca may be present. The shape and composition of the telson differs between arthropod groups. Crustaceans In lobsters, Caridea, shrimp and other Decapoda, decapods, the telson, along with the uropods, forms the tail fan. This is used as a paddle in the caridoid escape reaction ("lobstering"), whereby an alarmed animal rapidly flexes its tail, causing it to dart backwards. Krill can reach speeds of over 60 cm per second by this means. The Induction period, trigger time to optical stimulus (physiology), stimulus is, in spite of the low temperatures, only 55 milliseconds. In th ...
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Exopod
The arthropod leg is a form of jointed appendage of arthropods, usually used for walking. Many of the terms used for arthropod leg segments (called podomeres) are of Latin origin, and may be confused with terms for bones: ''coxa'' (meaning hip (anatomy), hip, : ''coxae''), ''trochanter'', ''femur'' (: ''femora''), ''tibia'' (: ''tibiae''), ''tarsus'' (: ''tarsi''), ''ischium'' (: ''ischia''), ''metatarsus'', ''carpus'', ''dactylus'' (meaning finger), ''patella'' (: ''patellae''). Homology (biology), Homologies of leg segments between groups are difficult to prove and are the source of much argument. Some authors posit up to eleven segments per leg for the most recent common ancestor of Neontology, extant arthropods but modern arthropods have eight or fewer. It has been argued that the ancestral leg need not have been so complex, and that other events, such as successive loss of function of a Homeobox, ''Hox''-gene, could result in Parallel evolution, parallel gains of leg segments ...
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Seta
In biology, setae (; seta ; ) are any of a number of different bristle- or hair-like structures on living organisms. Animal setae Protostomes Depending partly on their form and function, protostome setae may be called macrotrichia, chaetae, Scale (insect anatomy), scales, or Common name, informally, hairs. The setal membrane is not Cuticle, cuticularized, so movement is possible. Annelid setae are stiff bristles present on the body. They allow earthworms and their relatives to attach to the surface and prevent backsliding during peristaltic motion. These hairs make it difficult to pull a worm straight from the ground. Setae in oligochaetes (the group including earthworms) are largely composed of chitin. They are classified according to the limb to which they are attached; for instance, notosetae are attached to notopodia; neurosetae to neuropodia. The setae on polychaete worms are referred to as chaeta due to their differing morphology. Crustaceans have mechano- and chemos ...
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Hypostome (trilobite)
The hypostome is the hard mouthpart of trilobites found on the ventral side of the cephalon (head). The hypostome can be classified into three types based on whether they are permanently attached to the rostrum or not and whether they are aligned to the anterior dorsal tip of the glabella. Morphology The center of the hypostome is an ovoid, typically convex part called the median body, often divided into an anterior lobe and a posterior lobe. Either side of the median body is a border with various extensions, including anterior and posterior wings, sometimes bearing knob-like processes. The hypostome is hollow, and encloses the mouthparts, the anterior digestive tract, and the bases of the antennae. Trilobite antennae pass through notches between the anterior and posterior wings, then forward. The anterior wings are designed to rest firmly against internal structures (ventral apodemes) on the glabella. Variation in trilobite hypostome morphology is crucial in modern discussi ...
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Genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In binomial nomenclature, the genus name forms the first part of the binomial species name for each species within the genus. :E.g. ''Panthera leo'' (lion) and ''Panthera onca'' (jaguar) are two species within the genus ''Panthera''. ''Panthera'' is a genus within the family Felidae. The composition of a genus is determined by taxonomy (biology), taxonomists. The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including the idea that a newly defined genus should fulfill these three criteria to be descriptively useful: # monophyly – all descendants of an ancestral taxon are grouped together (i.e. Phylogeneti ...
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