Lithocalla Ecorticata
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Lithocalla Ecorticata
''Lithocalla'' is a small genus of lichen-forming fungi in the family Ramalinaceae. It comprises two known species that form crustose lichen, crust-like growths with a powdery texture on rocks. The genus was established in 2020 based on genetic analysis of species previously classified under different genera. ''Lithocalla'' lichens are characterised by their pale yellow-green colour and their preference for sheltered, shaded rock surfaces. They lack visible reproductive structures and likely spread through fragmentation. The two species have distinct geographical distributions: ''Lithocalla ecorticata, L. ecorticata'' is found mainly in Great Britain and Norway, while ''Lithocalla malouina, L. malouina'' is native to the Falkland Islands. Taxonomy The genus Lithocalla was erected in 2020 by Alan Orange to accommodate two species of leprose lichens containing usnic acid. It is placed in the family Ramalinaceae, order Lecanorales, though its exact phylogenetic position wi ...
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Alan Orange
Alan Orange (19 June 1955 – 5 February 2023) was a British lichenologist. His research interests included lichen taxonomy and phylogenetics, aquatic lichens, and the family Verrucariaceae. Early life and education Alan Orange was born in Leeds on 19 June 1955, and shortly after, his family relocated to Longhope in the Forest of Dean. This environment fostered his passion for botanical exploration, often accompanied by cousins and the family dogs. His penchant for collecting and classifying challenging organisms began in childhood, evident from his extensive bramble collection. Additionally, he showcased a natural talent for art, crafting intricate botanical illustrations throughout his life. Pursuing his passion academically, he completed his undergraduate studies at Bristol University, followed by a master's degree in pure and applied plant taxonomy at Reading University, earning it with distinction. Career and contributions In the early 1980s, Orange worked as a field ...
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Molecular Phylogenetic
Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to determine the processes by which diversity among species has been achieved. The result of a molecular phylogenetic analysis is expressed in a phylogenetic tree. Molecular phylogenetics is one aspect of molecular systematics, a broader term that also includes the use of molecular data in taxonomy and biogeography. Molecular phylogenetics and molecular evolution correlate. Molecular evolution is the process of selective changes (mutations) at a molecular level (genes, proteins, etc.) throughout various branches in the tree of life (evolution). Molecular phylogenetics makes inferences of the evolutionary relationships that arise due to molecular evolution and results in the construction of a phylogenetic tree. History The theoretical fra ...
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Lichen Genera
A lichen ( , ) is a hybrid colony (biology), colony of algae or cyanobacteria living symbiotically among hypha, filaments of multiple fungus species, along with yeasts and bacteria embedded in the cortex or "skin", in a mutualism (biology), mutualistic relationship.Introduction to Lichens – An Alliance between Kingdoms
. University of California Museum of Paleontology. .
Lichens are the lifeform that first brought the term symbiosis (as ''Symbiotismus'') into biological context. Lichens have since been recognized as important actors in nutrient cycling and producers which many higher trophic feeders feed on, such as reindeer, gastropods, nematodes, mites, and springtails. Lichens have properties different from those of their component organisms. They come in man ...
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Lecanorales Genera
The Lecanorales are an order of mostly lichen-forming fungi belonging to the class Lecanoromycetes in the division Ascomycota. The order contains 26 families, 269 genera, and 5695 species. Families Suborder Lecanorineae * Biatorellaceae M. Choisy ex Hafellner & Casares-Porcel, 1992 * Brigantiaeaceae Hafellner & Bellem., 1982 * Bruceomycetaceae Rikkinen & A.R.Schmidt in Rikkinen et al., * Byssolomataceae Zahlbr. 1926 * Carbonicolaceae Bendiksby & Timdal (2013) * Catillariaceae Hafellner, 1984 * Cetradoniaceae J.C. Wei & Ahti 2002 * Cladoniaceae Zenker, J.C. 1827–1829 * Dactylosporaceae Bellem. & Hafellner, 1982 * Gypsoplacaceae Timdal, E. 1990 * Haematommataceae Hafellner, 1984 * Lecanoraceae Fée, A.L.A. 1824 * Malmideaceae Kalb, K., Rivas Plata, E., Lücking, R. & Lumbsch, H.T. 2011 * Pachyascaceae Poelt ex P.M.Kirk, P.F.Cannon & J.C.David, 2001 * Parmeliaceae The Parmeliaceae is a large and diverse family of Lecanoromycetes. With over 2700 species i ...
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Overhang (rock Formation)
In rock climbing, an overhang is a type of route that leans back at an angle of over 90 degrees for part or all of the climb, and at its most severe can be a horizontal roof. Overhang (and roof) climbs have existed throughout climbing, originally in aid climbing where mechanical devices were used to first scale them. They became more common in free climbing during the 1990s as sport climbers used new training methods that enabled them to climb routes that were continuously, and severely, overhanging. Overhangs (and roofs) also feature prominently in advanced bouldering and in competition climbing. Overhanging routes require a range of techniques to manage the demands placed on the upper body, as the feet are less weighted. Body positioning — keeping the hips close to the rock face and the arms straight — is important. Advanced techniques such as the drop-knee, side-pulls-underclings-gastons, and the full range of heel and toe hooks are used for this. Knee-bars and bat ...
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Endemic
Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsewhere. For example, the Cape sugarbird is found exclusively in southwestern South Africa and is therefore said to be ''endemic'' to that particular part of the world. An endemic species can also be referred to as an ''endemism'' or, in scientific literature, as an ''endemite''. Similarly, many species found in the Western ghats of India are examples of endemism. Endemism is an important concept in conservation biology for measuring biodiversity in a particular place and evaluating the risk of extinction for species. Endemism is also of interest in evolutionary biology, because it provides clues about how changes in the environment cause species to undergo range shifts (potentially expanding their range into a larger area or bec ...
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Drystone
Dry stone, sometimes called drystack or, in Scotland, drystane, is a building method by which structures are constructed from stones without any mortar to bind them together. A certain amount of binding is obtained through the use of carefully selected interlocking stones. Dry stone construction is best known in the context of stone walls, traditionally used for the boundaries of fields and churchyards, or as retaining walls for terracing, but dry stone shelters, houses and other structures also exist. The term tends not to be used for the many historic styles which used precisely-shaped stone, but did not use mortar, for example the Greek temple and Inca architecture. The art of dry stone walling was inscribed in 2018 on the UNESCO representative list of the intangible cultural heritage of humanity, for dry stone walls in countries such as France, Greece, Italy, Slovenia, Croatia, Switzerland and Spain. In 2024, Ireland was added to the list. History Some dry stone wa ...
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Siliceous
Silicon dioxide, also known as silica, is an oxide of silicon with the chemical formula , commonly found in nature as quartz. In many parts of the world, silica is the major constituent of sand. Silica is one of the most complex and abundant families of materials, existing as a compound of several minerals and as a synthetic product. Examples include fused quartz, fumed silica, opal, and aerogels. It is used in structural materials, microelectronics, and as components in the food and pharmaceutical industries. All forms are white or colorless, although impure samples can be colored. Silicon dioxide is a common fundamental constituent of glass. Structure In the majority of silicon dioxides, the silicon atom shows tetrahedral coordination, with four oxygen atoms surrounding a central Si atomsee 3-D Unit Cell. Thus, SiO2 forms 3-dimensional network solids in which each silicon atom is covalently bonded in a tetrahedral manner to 4 oxygen atoms. In contrast, CO2 is a linear ...
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Green Alga
The green algae (: green alga) are a group of chlorophyll-containing autotrophic eukaryotes consisting of the phylum Prasinodermophyta and its unnamed sister group that contains the Chlorophyta and Charophyta/ Streptophyta. The land plants ( Embryophytes) have emerged deep within the charophytes as a sister of the Zygnematophyceae. Since the realization that the Embryophytes emerged within the green algae, some authors are starting to include them. The completed clade that includes both green algae and embryophytes is monophyletic and is referred to as the clade Viridiplantae and as the kingdom Plantae. The green algae include unicellular and colonial flagellates, most with two flagella per cell, as well as various colonial, coccoid (spherical), and filamentous forms, and macroscopic, multicellular seaweeds. There are about 22,000 species of green algae, many of which live most of their lives as single cells, while other species form coenobia (colonies), long filaments, or ...
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Medulla (lichenology)
The medulla is a horizontal layer within a lichen thallus. It is a loosely arranged layer of interlaced hyphae below the upper cortex and photobiont A lichen ( , ) is a hybrid colony of algae or cyanobacteria living symbiotically among filaments of multiple fungus species, along with yeasts and bacteria embedded in the cortex or "skin", in a mutualistic relationship.
zone, but above the lower cortex.Galloway, D.J. (1992). Flora of Australia - ''Lichen Glossary'' The medulla generally has a cottony appearance. It is the widest layer of a heteromerous lichen thallus.


References

Fungal morphology and anatomy Lichenology {{lichen-stub ...
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Hypha
A hypha (; ) is a long, branching, filamentous structure of a fungus, oomycete, or actinobacterium. In most fungi, hyphae are the main mode of vegetative growth, and are collectively called a mycelium. Structure A hypha consists of one or more cells surrounded by a tubular cell wall. In most fungi, hyphae are divided into cells by internal cross-walls called "septa" (singular septum). Septa are usually perforated by pores large enough for ribosomes, mitochondria, and sometimes nuclei to flow between cells. The major structural polymer in fungal cell walls is typically chitin, in contrast to plants and oomycetes that have cellulosic cell walls. Some fungi have aseptate hyphae, meaning their hyphae are not partitioned by septa. Hyphae have an average diameter of 4–6 μm. Growth Hyphae grow at their tips. During tip growth, cell walls are extended by the external assembly and polymerization of cell wall components, and the internal production of new cell membrane. ...
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Thallus
Thallus (: thalli), from Latinized Greek (), meaning "a green shoot" or "twig", is the vegetative tissue of some organisms in diverse groups such as algae, fungi, some liverworts, lichens, and the Myxogastria. A thallus usually names the entire body of a multicellular non-moving organism in which there is no organization of the tissues into organs. Many of these organisms were previously known as the thallophytes, a polyphyletic group of distantly related organisms. An organism or structure resembling a thallus is called thalloid, thalloidal, thalliform, thalline, or thallose. Even though thalli do not have organized and distinct parts ( leaves, roots, and stems) as do the vascular plants, they may have analogous structures that resemble their vascular "equivalents". The analogous structures have similar function or macroscopic structure, but different microscopic structure; for example, no thallus has vascular tissue. In exceptional cases such as the Lemnoideae, where th ...
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