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Hormogoneae
Hormogonia are motile filaments of cells formed by some cyanobacteria in the order Nostocales and Stigonematales. They are formed during vegetative reproduction in unicellular, filamentous cyanobacteria, and some may contain heterocysts and akinetes. Cyanobacteria differentiate into hormogonia when exposed to an environmental stress or when placed in new media. Hormogonium differentiation is crucial for the development of nitrogen-fixing plant cyanobacteria symbioses, in particular that between cyanobacteria of the genus ''Nostoc'' and their hosts. In response to a hormogonium-inducing factor (HIF) secreted by plant hosts, cyanobacterial symbionts differentiate into hormogonia and then dedifferentiate back into vegetative cells after about 96 hours. Hopefully, they have managed to reach the plant host by this time. The bacteria then differentiate specialized nitrogen-fixing cells called heterocysts and enter into a working symbiosis with the plant. Depending on species, Hormogo ...
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Nostoc With Hormogonia 40x
''Nostoc'', also known as star jelly, troll's butter, spit of moon, fallen star, witch's butter (not to be confused with the fungi commonly known as witches' butter), and witch's jelly, is the most common genus of cyanobacteria found in a variety of both aquatic and terrestrial environments that may form colonies composed of filaments of moniliform cells in a gelatinous sheath of polysaccharides. It may also grow symbiotically within the tissues of plants, providing nitrogen to its host through the action of terminally differentiated cells known as heterocysts. ''Nostoc'' is a genus that includes many species that are diverse in morphology, habitat distribution, and ecological function. ''Nostoc'' can be found in soil, on moist rocks, at the bottom of lakes and springs, and rarely in marine habitats. It may also be found in terrestrial temperate, desert, tropical, or polar environments. The name ''Nostoc'' was coined by Paracelsus and is a combination of the English ''nost''ril ...
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Cyanobacteria
Cyanobacteria ( ) are a group of autotrophic gram-negative bacteria that can obtain biological energy via oxygenic photosynthesis. The name "cyanobacteria" () refers to their bluish green (cyan) color, which forms the basis of cyanobacteria's informal common name, blue-green algae. Cyanobacteria are probably the most numerous taxon to have ever existed on Earth and the first organisms known to have produced oxygen, having appeared in the middle Archean eon and apparently originated in a freshwater or terrestrial environment. Their photopigments can absorb the red- and blue-spectrum frequencies of sunlight (thus reflecting a greenish color) to split water molecules into hydrogen ions and oxygen. The hydrogen ions are used to react with carbon dioxide to produce complex organic compounds such as carbohydrates (a process known as carbon fixation), and the oxygen is released as a byproduct. By continuously producing and releasing oxygen over billions of years, cyanobacte ...
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Nostocales
The Nostocales are an order of cyanobacteria Cyanobacteria ( ) are a group of autotrophic gram-negative bacteria that can obtain biological energy via oxygenic photosynthesis. The name "cyanobacteria" () refers to their bluish green (cyan) color, which forms the basis of cyanobacteri ... containing most of its species. It includes filamentous forms, both simple or branched, and both those occurring as single strands or multiple strands within a sheath. Some members show a decrease in width from the base, and some have heterocysts. References Bacteria orders {{cyanobacteria-stub ...
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Stigonematales
The Rivulariaceae are a family of cyanobacteria within the Nostocales in which the filaments (trichomes) are tapered from wider at the base to narrower at the tip. The type species is '' Rivularia haematites'' (Dc) C. A. Agardh. Genera , WoRMS listed the following genera under Rivulariaceae: * '' Amphithrix'' Bornet & Flahault, 1886 * '' Ariasmontanoa'' Molinari & Guiry, 2021 * '' Calothrix'' C.Agardh ex Bornet & Flahault, 1886 * '' Dichothrix'' Zanardini ex Bornet & Flahault, 1886 * '' Gaillardotella'' Bory de Saint-Vincent ex Kuntze, 1898 * '' Gardnerula'' G.De Toni, 1936 * '' Gloiotrichia'' J.Agardh, 1842 * '' Heteractis'' A.P.de Candolle, 1838 * '' Isactis'' Thuret ex Bornet & Flahault, 1886 * '' Macrochaete'' Berrendero, J.R.Johansen & Kastovsky, 2016 * '' Mastigonema'' H.Schwabe ex A.B.Frank, 1886 * '' Montanoa'' P.González, 1947 * '' Nunduva'' L.González-Resendiz, H.León-Tejera & J.R.Johansen, 2018 * '' Phyllonema'' D.O.Alvarenga, J.Rigonato, L.H.Z.Branco, I.S.Melo ...
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Heterocyst
Heterocysts or heterocytes are specialized nitrogen-fixing cells formed during nitrogen starvation by some filamentous cyanobacteria, such as ''Nostoc'', ''Cylindrospermum'', and '' Anabaena''. They fix nitrogen from dinitrogen (N2) in the air using the enzyme nitrogenase, in order to provide the cells in the filament with nitrogen for biosynthesis. Nitrogenase is inactivated by oxygen, so the heterocyst must create a microanaerobic environment. The heterocysts' unique structure and physiology require a global change in gene expression. For example, heterocysts: * produce three additional cell walls, including one of glycolipid that forms a hydrophobic barrier to oxygen * produce nitrogenase and other proteins involved in nitrogen fixation * degrade photosystem II, which produces oxygen * up-regulate glycolytic enzymes * produce proteins that scavenge any remaining oxygen * contain polar plugs composed of cyanophycin which slows down cell-to-cell diffusion Cyanobacteria usu ...
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Akinete
An akinete is an enveloped, thick-walled, non-motile, dormant cell (biology), cell formed by both cyanobacteria and algae. Cyanobacterial akinetes are mainly formed by filamentous, heterocyst-forming members under the order Nostocales and Stigonematales. Eukaryotic microalgae also produce akinetes, such as ''Haematococcus''. During akinete formation, cells accumulate and store various essential material, allowing the akinete to serve as a survival structure for up to many years. The cell akinetes are resistant to cold and desiccation. However, akinetes are not resistant to heat. Once conditions become more favorable for growth, the akinete can then germinate back into a vegetative cell. Increased light intensity, nutrients availability, oxygen availability, and changes in salinity are important triggers for germination. In cyanobacteria, akinetes usually develop in strings with each cell differentiating after another and this occurs next to heterocysts if they are present. Develo ...
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Stress (biology)
Stress, whether physiological, biological or psychological, is an organism's response to a stressor, such as an environmental condition or change in life circumstances. When stressed by stimuli that alter an organism's environment, multiple systems respond across the body. In humans and most mammals, the autonomic nervous system and Hypothalamic–pituitary–adrenal axis, hypothalamic-pituitary-adrenal (HPA) axis are the two major systems that respond to stress. Two well-known hormones that humans produce during stressful situations are adrenaline and cortisol. The Sympathoadrenal system, sympathoadrenal medullary axis (SAM) may activate the fight-or-flight response through the sympathetic nervous system, which dedicates energy to more relevant bodily systems to Acute stress reaction, acute adaptation to stress, while the parasympathetic nervous system returns the body to homeostasis. The second major physiological stress-response center, the HPA axis, regulates the release ...
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Nitrogen Fixation
Nitrogen fixation is a chemical process by which molecular dinitrogen () is converted into ammonia (). It occurs both biologically and abiological nitrogen fixation, abiologically in chemical industry, chemical industries. Biological nitrogen fixation or ''diazotrophy'' is catalyzed by enzymes called nitrogenases. These enzyme complexes are encoded by the Nif gene, ''Nif'' genes (or ''Nif'' homologs) and contain iron, often with a second metal (usually molybdenum, but sometimes vanadium). Some nitrogen-fixing bacteria have symbiotic relationships with plants, especially legumes, mosses and aquatic ferns such as ''Azolla''. Looser non-symbiotic relationships between diazotrophs and plants are often referred to as associative, as seen in nitrogen fixation on rice roots. Nitrogen fixation occurs between some termites and fungus, fungi. It occurs naturally in the air by means of NOx, NOx production by lightning. Fixed nitrogen is essential to life on Earth. Organic compounds such ...
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Nostoc
''Nostoc'', also known as star jelly, troll's butter, spit of moon, fallen star, witch's butter (not to be confused with the fungi commonly known as witches' butter), and witch's jelly, is the most common genus of cyanobacteria found in a variety of both aquatic and terrestrial environments that may form colony (biology), colonies composed of Filamentation, filaments of wiktionary:moniliform, moniliform cells in a gelatinous sheath of polysaccharides. It may also grow symbiosis, symbiotically within the tissue (biology), tissues of plants, providing nitrogen to its host through the action of terminally differentiated cells known as heterocysts. ''Nostoc'' is a genus that includes many species that are diverse in morphology, habitat distribution, and ecological function. ''Nostoc'' can be found in soil, on moist rocks, at the bottom of lakes and springs, and rarely in marine habitats. It may also be found in terrestrial temperate, desert, tropical, or polar environments. The name ' ...
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Host (biology)
In biology and medicine, a host is a larger organism that harbours a smaller organism; whether a parasite, parasitic, a mutualism (biology), mutualistic, or a commensalism, commensalist ''guest'' (symbiont). The guest is typically provided with nourishment and shelter. Examples include animals playing host to parasitic worms (e.g. nematodes), cell (biology), cells harbouring pathogenic (disease-causing) viruses, or a Fabaceae, bean plant hosting mutualistic (helpful) Rhizobia, nitrogen-fixing bacteria. More specifically in botany, a host plant supplies nutrient, food resources to micropredators, which have an evolutionarily stable strategy, evolutionarily stable relationship with their hosts similar to ectoparasitism. The host range is the collection of hosts that an organism can use as a partner. Symbiosis Symbiosis spans a wide variety of possible relationships between organisms, differing in their permanence and their effects on the two parties. If one of the partners in an ...
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Bacterial Gliding
Gliding motility is a type of translocation used by microorganisms that is independent of propulsive structures such as flagella, pili, and fimbriae. Gliding allows microorganisms to travel along the surface of low aqueous films. The mechanisms of this motility are only partially known. Twitching motility also allows microorganisms to travel along a surface, but this type of movement is jerky and uses pili as its means of transport. Bacterial gliding is a type of gliding motility that can also use pili for propulsion. The speed of gliding varies between organisms, and the reversal of direction is seemingly regulated by some sort of internal clock. For example the apicomplexans are able to travel at fast rates between 1–10 μm/s. In contrast '' Myxococcus xanthus'' bacteria glide at a rate of 0.08 μm/s. Types of motility Bacterial gliding is a process of motility whereby a bacterium can move under its own power. Generally, the process occurs whereby the bacteriu ...
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