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Hilomonadea
Rigifilida is a clade of non-ciliate phagotrophic eukaryotes. It consists of two genera: '' Micronuclearia'' and '' Rigifila''. Characteristics Cells of rigifilids are covered with either a single or a double-layered submembrane pellicular lamina that makes them rigid in consistence. Slender branching filopodia emanate from a ventral aperture of the cell and are employed to collect bacteria upon which they feed and to attach the organism to the substratum. Around this aperture, the pellicle is reflexed around forming a peristomial collar. Other notable features are flat and irregular shaped mitocondrial cristae, a single dorsal nucleus and the lack of centrioles and cilia. Phylogeny Taxonomy Rigifilida is currently placed in CRuMs. *Order Rigifilida Cavalier-Smith 2012 icronucleariida Cavalier-Smith 2008** Family Rigifilidae ''Rigifila'' () is a genus of free-living single-celled eukaryotes, or protists, containing the sole species ''Rigifila ramosa'' (). It is classifie ...
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Rigifila Ramosa
''Rigifila'' () is a genus of free-living single-celled eukaryotes, or protists, containing the sole species ''Rigifila ramosa'' (). It is classified within the monotypic family Rigifilidae. Along with Micronucleariidae, it is a member of Rigifilida, an order of basal eukaryotes within the CRuMs clade. It differs from ''Micronuclearia'' by having two proteic layers surrounding their cytoplasm instead of a single one, and having more irregular mitochondrial cristae, among other morphological differences. Systematics Etymology The name of this genus is derived , referring to the rigid the cell body due to their submembrane proteic layers, and ''filum'' 'thread', referring to the filopodia (thin thread-like pseudopodia). The species epithet is derived , referring to the branched filopodia exhibited by the organism. Classification The species ''Rigifila ramosa'' was described in 2012 by biologists Akinori Yabuki, Ken-Ichiro Ishida and Thomas Cavalier-Smith from freshwater ...
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Collodictyonidae
Collodictyonidae (also Diphylleidae) is a group of aquatic, unicellular eukaryotic organisms with two to four terminal flagella. They feed by phagocytosis, ingesting other unicellular organisms like algae and bacteria Bacteria (; : bacterium) are ubiquitous, mostly free-living organisms often consisting of one Cell (biology), biological cell. They constitute a large domain (biology), domain of Prokaryote, prokaryotic microorganisms. Typically a few micr .... Recent molecular analyses place collodictyonids (e.g. '' Collodictyon'') in a clade also containing Rigifilida and Mantamonadidae. This clade has been named CRuMs and is sister to Amorphea. Phylogeny Taxonomy * Class Diphyllatea Cavalier-Smith 2003 nisomonadea ; Diphyllatia Cavalier-Smith 2003** Order Diphylleida Cavalier-Smith 1993 ollodictyonida *** Family Sulcomonadidae Cavalier-Smith 2012 **** Genus '' Sulcomonas'' Brugerolle 2006 ***** Species '' S. lacustris'' Brugerolle 2006 *** Family Collodictyon ...
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CRuMs
CRuMs is a clade of microbial eukaryotes, whose name is an acronym of the following constituent groups: i) Diphylleids, ii) rigifilids and iii) mantamonads as sister of the Amorphea. A new CRuMs order Glissandrida was proposed in 2025 to place the yet unclassified genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ... '' Glissandra''. It more or less supersedes Varisulca, as Ancyromonadida are inferred not to be specifically related to the orders Diphylleida / Collodictyonida, Rigifilida and Mantamonadida. Phylogeny References {{Taxonbar, from1=Q134939335, from2=Q59153571 Eukaryote taxa Podiata Protist taxa CRuMs taxa ...
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Apusomonadida
The apusomonads (family Apusomonadidae) are a group of protozoan zooflagellates that glide on surfaces, and mostly consume prokaryotes. They are of particular evolutionary interest because they appear to be the sister group to the Opisthokonts, the clade that includes both animals and fungi. Together with the Breviatea, these form the Obazoa clade. Characteristics Apusomonads are small gliding heterotrophic biflagellates (i.e. with two flagella) that possess a proboscis, formed partly or entirely by the anterior flagellum surrounded by a membranous sleeve. There is a pellicle under the dorsal cell membrane that extends into the proboscis sleeve and into a skirt that covers the sides of the cell. Apusomonads present two different cell plans: *Derived cell plan, represented by '' Apusomonas'', with a round cell body and a mastigophore, a projection of the cell containing both basal bodies at its end. *"''Amastigomonas''-like" cell plan, with an oval or oblong cell that gener ...
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Breviatea
Breviatea, commonly known as breviate amoebae, are a group of free-living, amitochondriate protists with uncertain phylogenetic position. They are biflagellate, and can live in anaerobic (oxygen-free) environments. They are currently placed in the Obazoa clade. They likely do not possess vinculin proteins. Their metabolism relies on fermentative production of ATP as an adaptation to their low-oxygen environment. The lineage emerged roughly one billion years ago, at a time when the oxygen content of the Earth's oceans was low, and they thus developed anaerobic lifestyles. Together with Apusomonads, they are the closest relatives of the Opisthokonts, a group that includes animals and fungi. Characteristics Mitochondrion-related organelles Mitochondrion-related organelles (MROs) are organelles that evolved from a degradation of ancestral, fully functional mitochondria. Among Breviatea, MROs are present in '' Pygsuia'', '' Breviata'' and '' Subulatomonas''. In the cells of '' ...
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Obazoa
Obazoa is a proposed sister clade of Amoebozoa (which together form Amorphea). The term Obazoa is based on the OBA acronym for Opisthokonta, Breviatea, and Apusomonadidae, the group's three constituent clades. Determining the placement of Breviatea and Apusomonadida and their properties is of interest for the development of the opisthokonts in which the main lineages of animals and fungi A fungus (: fungi , , , or ; or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and mold (fungus), molds, as well as the more familiar mushrooms. These organisms are classified as one ... emerged. The relationships among opisthokonts, breviates and apusomonads are not conclusively resolved (as of 2018), though Breviatea is usually inferred to be the most basal of the three lineages. The phylogeny of the Obazoa is shown in the cladogram. References {{Taxonbar, from=Q22087764 Amorphea taxa ...
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Amoebozoa
Amoebozoa is a major Taxonomy (biology), taxonomic group containing about 2,400 described species of Amoeba, amoeboid protists, often possessing blunt, fingerlike, Pseudopod#Morphology, lobose pseudopods and tubular mitochondrial cristae. In traditional classification schemes, Amoebozoa is usually ranked as a phylum within either the kingdom (biology), kingdom Protista or the kingdom Protozoa. In the classification favored by the International Society of Protistologists, it is retained as an unranked "supergroup (biology), supergroup" within Eukaryota. Molecular genetics, Molecular genetic analysis supports Amoebozoa as a monophyletic clade. Modern studies of eukaryotic phylogenetic trees identify it as the sister group to Opisthokonta, another major clade which contains both fungi and animals as well as several other clades comprising some 300 species of unicellular eukaryotes. Amoebozoa and Opisthokonta are sometimes grouped together in a high-level taxon, named Amorphea. Amoeboz ...
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