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Heterosteus Ingens
''Heterosteus'' (also known as ''Heterostius'') is an extinct genus of heterosteid placoderm of the Middle Devonian known from remains discovered in Europe and Greenland. Name ''Heterosteus'' was originally described in 1837 as species of '' Trionyx'', a softshelled turtle. Also in later studies it was often misidentified and given names like ''Ichthyosauroides'', '' Asterolepis asmussi'' and ''Chelonichthys asmusii''. Even in recent studies, it is controversial as to whether to use genus name ''Heterosteus'' or ''Heterostius''. According to International Code of Zoological Nomenclature, suffix ‘-ostius’ in scientific name should not be corrected as ‘-osteus’, so some study uses genus name ''Heterostius''. Description This genus includes the largest species in the family, and are among the largest arthrodires, as well, with the type species, ''H. asmussi'', having an estimated body length of up to . The genus differs from ''Herasmius'' by having the orbits on slight ...
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Middle Devonian
The Devonian ( ) is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, million years ago (Mya), to the beginning of the Carboniferous, Mya. It is named after Devon, England, where rocks from this period were first studied. The first significant adaptive radiation of life on dry land occurred during the Devonian. Free-sporing vascular plants began to spread across dry land, forming extensive forests which covered the continents. By the middle of the Devonian, several groups of plants had evolved leaves and true roots, and by the end of the period the first seed-bearing plants appeared. The arthropod groups of myriapods, arachnids and hexapods also became well-established early in this period, after starting their expansion to land at least from the Ordovician period. Fish reached substantial diversity during this time, leading the Devonian to often be dubbed the Age of Fishes. The placoderms began dominating al ...
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Heterosteus Ingens
''Heterosteus'' (also known as ''Heterostius'') is an extinct genus of heterosteid placoderm of the Middle Devonian known from remains discovered in Europe and Greenland. Name ''Heterosteus'' was originally described in 1837 as species of '' Trionyx'', a softshelled turtle. Also in later studies it was often misidentified and given names like ''Ichthyosauroides'', '' Asterolepis asmussi'' and ''Chelonichthys asmusii''. Even in recent studies, it is controversial as to whether to use genus name ''Heterosteus'' or ''Heterostius''. According to International Code of Zoological Nomenclature, suffix ‘-ostius’ in scientific name should not be corrected as ‘-osteus’, so some study uses genus name ''Heterostius''. Description This genus includes the largest species in the family, and are among the largest arthrodires, as well, with the type species, ''H. asmussi'', having an estimated body length of up to . The genus differs from ''Herasmius'' by having the orbits on slight ...
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Westralichthys Uwagedensis
''Westralichthys'' is an extinct monospecific genus of dunkleosteoid from the Late Devonian: Middle Famennian stage from Western Australia. Phylogeny ''Westralichthys'' belongs to the superfamily Dunkleosteoidea, related to the giant '' Dunkleosteus''. The phylogeny A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological spec ... of ''Westralichthys'' can be shown in the cladogram below: References Arthrodires {{placoderm-stub ...
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Golshanichthys Asiatica
''Golshanichthys'' is an extinct monospecific genus of dunkleosteid from the Late Devonian: Frasnian stage from Kerman, Iran. Phylogeny ''Golshanichthys'' belongs to the family Dunkleosteidae, closely related to the giant ''Dunkleosteus''. The phylogeny of ''Golshanichthys'' can be shown in the cladogram below: Alternatively, the subsequent 2016 Zhu ''et al.'' study using a larger morphological dataset recovered Panxiosteidae well outside of Dunkleosteoidea Dunkleosteoidea is an extinct superfamily of arthrodire placoderms that lived during the Devonian period. The gigantic apex predator '' Dunkleosteus terrelli'' is the best known member of this group. Phylogeny Eubrachythoraci is divided into ..., leaving the status of Dunkleosteidae as a clade grouping separate from Dunkleosteoidea in doubt, as shown in the cladogram below: References Arthrodires Placoderms of Asia {{placoderm-stub ...
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Kiangyousteus Yohii
''Kiangyousteus'' is an extinct monotypic genus of dunkleosteid from the Middle Devonian: Givetian aged Guanwu Formation in the Sichuan province of south-western China. The type species, ''Kiangyousteus yohii'', was the first known arthrodire from Asia. Etymology ''Kiangyousteus'' is named after Jiangyou (formerly ‘Kiangyou’) district of Sichuan Province, the location where the fossils were found in 1953 by Professor S. H. Yoh of Peking University. Phylogeny ''Kiangyousteus'' belongs to the family Dunkleosteidae. The phylogeny of ''Kiangyousteus'' can be shown in the cladogram below: Alternatively, the subsequent 2016 Zhu ''et al.'' study using a larger morphological dataset recovered Panxiosteidae well outside of Dunkleosteoidea Dunkleosteoidea is an extinct superfamily of arthrodire placoderms that lived during the Devonian period. The gigantic apex predator '' Dunkleosteus terrelli'' is the best known member of this group. Phylogeny Eubrachythoraci is divi ...
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Eastmanosteus Pustulosus
''Eastmanosteus'' ("Eastman's bone") is a fossil genus of dunkleosteid placoderms. It was closely related to the giant ''Dunkleosteus'', but differed from that genus in size, in possessing a distinctive tuberculated bone ornament, a differently shaped nuchal plate and a more zig-zagging course of the sutures of the skull roof.http://www.ivpp.cas.cn/cbw/gjzdwxb/xbwzxz/200812/W020090813371138329343.pdf Species of ''Eastmanosteus'' had powerful jaws with sharp cutting edges and were likely active predators. Fossils have been found in many parts of the world in marine sediments dating from the Middle to Late Devonian. They were medium-to-large fish, with specimens ''E. pustulosus'' and ''E. licharevi'' approaching a total length of 3 metres. Complete exoskeletons with soft-tissue traces of ''E. calliaspis'' from Australia make this one of the best known dunkleosteids. Phylogeny ''Eastmanosteus'' and its relative ''Dunkleosteus'' belong to the family Dunkleosteidae. The phylogeny of ...
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Cladogram
A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to descendants, nor does it show how much they have changed, so many differing evolutionary trees can be consistent with the same cladogram. A cladogram uses lines that branch off in different directions ending at a clade, a group of organisms with a last common ancestor. There are many shapes of cladograms but they all have lines that branch off from other lines. The lines can be traced back to where they branch off. These branching off points represent a hypothetical ancestor (not an actual entity) which can be inferred to exhibit the traits shared among the terminal taxa above it. This hypothetical ancestor might then provide clues about the order of evolution of various features, adaptation, and other evolutionary narratives about an ...
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Dunkleosteus
''Dunkleosteus'' is an extinct genus of large armored, jawed fishes that existed during the Late Devonian period, about 382–358 million years ago. It consists of ten species, some of which are among the largest placoderms to have ever lived: ''D. terrelli'', ''D. belgicus'', ''D. denisoni'', ''D. marsaisi'', ''D. magnificus'', ''D. missouriensis'', ''D. newberryi'', ''D. amblyodoratus'', and ''D. raveri''. The largest and most well known species is ''D. terrelli'', which grew up to long and in weight. ''Dunkleosteus'' could quickly open and close its jaw, like modern-day suction feeders, and had a bite force of at the tip and at the blade edge. Numerous fossils of the various species have been found in North America, Poland, Belgium, and Morocco. Etymology ''Dunkleosteus'' was named in 1956 to honour David Dunkle (1911–1982), former curator of vertebrate paleontology at the Cleveland Museum of Natural History. The genus name ''Dunkleosteus'' combines David Dunkle's ...
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Dunkleosteoidea
Dunkleosteoidea is an extinct superfamily of arthrodire placoderms that lived during the Devonian period. The gigantic apex predator '' Dunkleosteus terrelli'' is the best known member of this group. Phylogeny Eubrachythoraci is divided into the clades Coccosteomorphi and Pachyosteomorphi, the latter of which can be further sub-divided into Aspinothoracidi and Dunkleosteoidea. Dunkleosteoidea was then considered to consist of the two sister families Dunkleosteidae and Panxiosteidae. However, the 2016 Zhu ''et al.'' phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ... study using a larger morphological dataset recovered Panxiosteidae well outside of Dunkleosteoidea, leaving the status of Dunkleosteidae as a clade grouping separate from Dunkleosteoidea in doubt ...
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Superfamily (biology)
In biological classification, taxonomic rank is the relative level of a group of organisms (a taxon) in an ancestral or hereditary hierarchy. A common system consists of species, genus, family, order, class, phylum, kingdom, domain. While older approaches to taxonomic classification were phenomenological, forming groups on the basis of similarities in appearance, organic structure and behaviour, methods based on genetic analysis have opened the road to cladistics. A given rank subsumes under it less general categories, that is, more specific descriptions of life forms. Above it, each rank is classified within more general categories of organisms and groups of organisms related to each other through inheritance of traits or features from common ancestors. The rank of any ''species'' and the description of its ''genus'' is ''basic''; which means that to identify a particular organism, it is usually not necessary to specify ranks other than these first two. Consider a particul ...
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Heterosteidae
Heterosteidae (also known as Heterostiidae) is an extinct family of moderately large to giant, flattened, benthic arthrodire placoderms with distinctive, flattened, triangular skulls that are extremely broad posteriorly, but become very narrow anteriorly. Heterosteidae belongs to the superfamily SUPERFAMILY is a database and search platform of structural and functional annotation for all proteins and genomes. It classifies amino acid sequences into known structural domains, especially into SCOP superfamilies. Domains are functional, str ... Dunkleosteoidea, a relative of the giant '' Dunkleosteus'', as shown in the cladogram below: Genera '' Herasmius'' Orvig, 1969 '' Heterosteus'' Asmuss, 1856 '' Yinostius'' J. Wang & N. Wang, 1984 References {{Taxonbar, from=Q15259423 Arthrodires Placoderm families Early Devonian first appearances Middle Devonian extinctions ...
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