Heterogyna
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Heterogyna
Heterogynaidae is a minor and disputed lineage (only eight described species in a single genus, ''Heterogyna'') of small spheciform wasps occurring in Madagascar, Botswana, Turkmenistan, Oman, the United Arab Emirates, and the Eastern Mediterranean area. The majority are dark in color and range in size from approximately 1.5 to 5.0 mm. Most specimens have been collected in arid climates, but one species from Madagascar is known to occur in a humid forest habitat. Although males have functional wings, heterogynaid females are typically brachypterous, a trait which is unique among spheciform wasps. Wing venation is reduced in both sexes. All species are diurnal, with the exception of ''H. nocticola''. Other aspects of their biology are completely unknown, but details of their morphology have prompted researchers to hypothesize that they may be non-fossorial parasitoids adapted to hunt in tight spaces, such as under tree bark. This is speculative and has not yet been confirmed ...
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Apoidea
The superfamily Apoidea is a major group (of over 30 000 species) within the Hymenoptera, which includes two traditionally recognized lineages, the "sphecoid" wasps, and the bees. Molecular phylogeny demonstrates that the bees arose from within the traditional " Crabronidae", so that grouping is paraphyletic, and this has led to a reclassification to produce monophyletic families.Manuela Sann, Oliver Niehuis, Ralph S. Peters, Christoph Mayer, Alexey Kozlov, Lars Podsiadlowski, Sarah Bank, Karen Meusemann, Bernhard Misof, Christoph Bleidorn and Michael Ohl (2018) Phylogenomic analysis of Apoidea sheds new light on the sister group of bees. ''BMC Evolutionary Biology'' 18:71. doi:10.1186/s12862-018-1155-8 Diagnostic features Apoid wasps and bees have several traits in common:O'Neill, K.M. (2008). Apoid Wasps (Hymenoptera: Apoidea: Spheciformes). In: Capinera, J.L. (eds) Encyclopedia of Entomology. Springer, Dordrecht. https://doi.org/10.1007/978-1-4020-6359-6_10300 * The ...
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Crabronidae
The Crabronidae is a large family of wasps within the superfamily Apoidea. Taxonomy and phylogeny This family has historically been treated as a subfamily in the now-defunct Spheciformes group under the family Sphecidae. The Spheciformes included well over 200 genera, containing well over 9000 species. Revision of these taxa resulted resulted in the restriction of the Sphecidae to what was once the subfamily Sphecinae. As a result, the former Crabroninae was elevated to family status as Crabronidae. Subsequent revision has further restricted the Crabronidae. Several of the subfamilies of the Crabronidae are often treated as families in their own right, as is true of the most recent phylogenies. Of these lineages of Apoidea, only three were not included within Crabronidae in the past: Ampulicidae, Sphecidae, and Anthophila. The following phylogenetic tree is based on Sann ''et al.'', 2018, which used phylogenomics to demonstrate that both the bees ( Anthophila) and the Sphe ...
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Fossorial
A fossorial animal () is one that is adapted to digging and which lives primarily (but not solely) underground. Examples of fossorial vertebrates are Mole (animal), moles, badgers, naked mole-rats, meerkats, armadillos, wombats, and mole salamanders. Among invertebrates, many molluscs (e.g., clams), insects (e.g., beetles, wasps, bees), and arachnids (e.g. spiders) are fossorial. Prehistoric evidence The physical adaptation of fossoriality is widely accepted as being widespread among many Prehistory, prehistoric Phylum, phyla and Taxon, taxa, such as bacteria and early eukaryotes. Furthermore, fossoriality has evolved independently multiple times, even within a single Family (biology), family. Fossorial animals appeared simultaneously with the colonization of land by arthropods in the late Ordovician period (over 440 million years ago). Other notable early burrowers include ''Eocaecilia'' and possibly ''Dinilysia''. The oldest example of burrowing in synapsids, the lineag ...
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Monophyly
In biological cladistics for the classification of organisms, monophyly is the condition of a taxonomic grouping being a clade – that is, a grouping of organisms which meets these criteria: # the grouping contains its own most recent common ancestor (or more precisely an ancestral population), i.e. excludes non-descendants of that common ancestor # the grouping contains all the descendants of that common ancestor, without exception Monophyly is contrasted with paraphyly and polyphyly as shown in the second diagram. A ''paraphyletic'' grouping meets 1. but not 2., thus consisting of the descendants of a common ancestor, excepting one or more monophyletic subgroups. A ''polyphyletic'' grouping meets neither criterion, and instead serves to characterize convergent relationships of biological features rather than genetic relationships – for example, night-active primates, fruit trees, or aquatic insects. As such, these characteristic features of a polyphyletic grouping are ...
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