Hadrosteus
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Hadrosteus
''Hadrosteus'' is an extinct monospecific genus of large arthrodire placoderm from the Late Frasnian (Late Devonian) Kellwasserkalk facies of Bad Wildungen, Germany. It had large, double-pronged inferognathals (lower jawbones), and serrated edges along its mandible, strongly suggesting that it was a fish-eating predator. The head had a triangular snout, and the trunkshield was short, but high, with a median dorsal plate that was broader than wide. The average skull length is about 16 centimeters. Etymology The type species ''Hadrosteus rapax'' means "Rapacious Strong-Bone". Phylogeny ''Hadrosteus'' is a member of the clade Aspinothoracidi, which belongs to the clade Pachyosteomorphi, one of the two major clades within Eubrachythoraci. The cladogram below shows the phylogeny A phylogenetic tree or phylogeny is a graphical representation which shows the evolutionary history between a set of species or Taxon, taxa during a specific time.Felsenstein J. (2004). ''Inferring Phylog ...
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Aspinothoracidi
Aspinothoracidi is a clade of placoderms, extinct armored fish most diverse during the Devonian. The gigantic apex predator ''Dinichthys'', is the best-known member of this group. Many other genera, such as the infamous ''Dunkleosteus'', were previously thought to be close relatives of ''Dinichthys'' and were grouped together in the family Dinichthyidae, though more recent studies have restricted that family to only its type species. Phylogeny Eubrachythoraci is divided into the clades Coccosteomorphi and Pachyosteomorphi, the latter of which can be further sub-divided into Aspinothoracidi and Dunkleosteoidea, as shown in the cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ... below: References Arthrodires {{Placoderm-stub ...
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Dunkleosteus Raveri
''Dunkleosteus'' is an extinct genus of large arthrodire ("jointed-neck") fish that existed during the Late Devonian period, about 382–358 million years ago. It was a pelagic fish inhabiting open waters, and one of the first vertebrate apex predators of any ecosystem. ''Dunkleosteus'' consists of ten species, some of which are among the largest placoderms ("plate-skinned") to have ever lived: ''D. terrelli'', ''D. belgicus'', ''D. denisoni'', ''D. marsaisi'', ''D. magnificus'', ''D. missouriensis'', ''D. newberryi'', ''D. amblyodoratus'', ''D. raveri'', and ''D. tuderensis.'' However, the validity of several of these species is unclear. The largest and best known species is ''D. terrelli''. Since body shape is not known, various methods of estimation put the living total length of the largest known specimen of ''D. terrelli'' between long and its weight around . Lengths of or more are poorly supported, with the most recent and extensive studies on the body shape and size of ' ...
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Dunkleosteus Terrelli
''Dunkleosteus'' is an extinct genus of large arthrodira, arthrodire ("jointed-neck") fish that existed during the Late Devonian period, about 382–358 million years ago. It was a pelagic fish inhabiting open waters, and one of the first vertebrate apex predators of any ecosystem. ''Dunkleosteus'' consists of ten species, some of which are among the largest placoderms ("plate-skinned") to have ever lived: ''D. terrelli'', ''D. belgicus'', ''D. denisoni'', ''D. marsaisi'', ''D. magnificus'', ''D. missouriensis'', ''D. newberryi'', ''D. amblyodoratus'', ''D. raveri'', and ''D. tuderensis.'' However, the validity of several of these species is unclear. The largest and best known species is ''D. terrelli''. Since body shape is not known, various methods of estimation put the living total length of the largest known specimen of ''D. terrelli'' between long and its weight around . Lengths of or more are poorly supported, with the most recent and extensive studies on the body shape a ...
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Pachyosteomorphi
Pachyosteomorphi is an extinct clade of arthrodire placoderms within the Eubrachythoraci (of the suborder Brachythoraci), armored fish most diverse during the Devonian. Most are considered to be pelagic (open ocean) long-distance swimmers, leading to their widespread distribution beginning from at least the Middle Devonian period. Phylogeny Pachyosteomorphi is the sister taxon to Coccosteomorphi, which together are the two main sub-clades of Eubrachythoraci. Pachyosteomorphi can be further sub-divided into Aspinothoracidi and Dunkleosteoidea, as shown in the cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ... below: References Arthrodires Devonian fish {{Placoderm-stub ...
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Eubrachythoraci
Eubrachythoraci is an extinct clade of arthrodire placoderms within the suborder Brachythoraci, armored fish most diverse during the Devonian. Most are considered to be pelagic (open ocean) long-distance swimmers, leading to their widespread distribution beginning from at least the Middle Devonian period. Phylogeny Brachythoraci is divided into the large derived clade Eubrachythoraci and several basal groups: Buchanosteoidea, Homosteidae, and Holonematidae. (Although Holonematidae's membership in Brachythoraci is disputed.) Eubrachythoraci is then further divided into the sub-clades Coccosteomorphi and Pachyosteomorphi, the latter of which can be further sub-divided into Aspinothoracidi and Dunkleosteoidea, as shown in the cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it doe ...
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Dunkleosteoidea
Dunkleosteoidea is an extinct superfamily of arthrodire placoderms that lived during the Devonian period. The gigantic apex predator ''Dunkleosteus terrelli'' is the best known member of this group. Phylogeny Eubrachythoraci is divided into the clades Coccosteomorphi and Pachyosteomorphi, the latter of which can be further sub-divided into Aspinothoracidi and Dunkleosteoidea. Dunkleosteoidea was then considered to consist of the two sister families Dunkleosteidae and Panxiosteidae. However, the 2016 Zhu ''et al.'' phylogenetic study using a larger morphological dataset recovered Panxiosteidae well outside of Dunkleosteoidea, leaving the status of Dunkleosteidae Dunkleosteidae is an extinct family (biology), family of arthrodire placoderms that lived during the Devonian period. The gigantic apex predator ''Dunkleosteus terrelli'' is the best known member of this group. Phylogeny While members of Dunkleo ... as a clade grouping separate from Dunkleosteoidea in doubt, as show ...
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Rhachiosteus Pterygiatus
''Rhachiosteus'' is an extinct monospecific genus of arthrodire placoderm from the Middle to Late Devonian of Germany. It is known only from a single specimen, which may be a larval or juvenile form, as the skull of said specimen is only 19 millimetres long. Phylogeny ''Rhachiosteus'' is a basal member of the clade Pachyosteomorphi, the sister taxon to Coccosteomorphi, which together are the two main sub-clades of Eubrachythoraci. The cladogram below shows the phylogeny A phylogenetic tree or phylogeny is a graphical representation which shows the evolutionary history between a set of species or Taxon, taxa during a specific time.Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, M ... of ''Rhachiosteus'': References Arthrodires Placoderms of Europe Arthrodire genera {{Placoderm-stub ...
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Coccosteomorphi
Coccosteomorphi is an extinct clade of arthrodire placoderms within the Eubrachythoraci (of the suborder Brachythoraci), armored fish most diverse during the Devonian. Most are considered to be pelagic (open ocean) long-distance swimmers, leading to their widespread distribution beginning from at least the Middle Devonian period. Phylogeny Coccosteomorphi is the sister taxon to Pachyosteomorphi, which together are the two main sub-clades of Eubrachythoraci. Coccosteomorphi can be further sub-divided into Coccosteoidea and Incisoscutoidea, as shown in the cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ... below: References Arthrodires {{Placoderm-stub ...
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Species
A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), classification and a taxonomic rank of an organism, as well as a unit of biodiversity. Other ways of defining species include their karyotype, DNA sequence, morphology (biology), morphology, behaviour, or ecological niche. In addition, palaeontologists use the concept of the chronospecies since fossil reproduction cannot be examined. The most recent rigorous estimate for the total number of species of eukaryotes is between 8 and 8.7 million. About 14% of these had been described by 2011. All species (except viruses) are given a binomial nomenclature, two-part name, a "binomen". The first part of a binomen is the name of a genus to which the species belongs. The second part is called the specific name (zoology), specific name or the specific ...
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Westralichthys Uwagedensis
''Westralichthys'' is an extinct monospecific genus of dunkleosteoid from the Late Devonian: Middle Famennian stage from Western Australia. It is estimated to be long. Phylogeny ''Westralichthys'' belongs to the superfamily Dunkleosteoidea, related to the giant ''Dunkleosteus''. The phylogeny of ''Westralichthys'' can be shown in the cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ... below: References Arthrodires {{placoderm-stub ...
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Golshanichthys Asiatica
''Golshanichthys'' is an extinct monospecific genus of dunkleosteid from the Late Devonian: Frasnian stage from Kerman, Iran. Phylogeny ''Golshanichthys'' belongs to the family Dunkleosteidae, closely related to the giant ''Dunkleosteus''. The phylogeny of ''Golshanichthys'' can be shown in the cladogram below: Alternatively, the subsequent 2016 Zhu ''et al.'' study using a larger morphological dataset recovered Panxiosteidae well outside of Dunkleosteoidea, leaving the status of Dunkleosteidae Dunkleosteidae is an extinct family (biology), family of arthrodire placoderms that lived during the Devonian period. The gigantic apex predator ''Dunkleosteus terrelli'' is the best known member of this group. Phylogeny While members of Dunkleo ... as a clade grouping separate from Dunkleosteoidea in doubt, as shown in the cladogram below: References Arthrodires Placoderms of Asia {{placoderm-stub ...
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Kiangyousteus Yohii
''Kiangyousteus'' is an extinct monotypic genus of dunkleosteid from the Middle Devonian: Givetian aged Guanwu Formation in the Sichuan province of south-western China. The type species, ''Kiangyousteus yohii'', was the first known arthrodire from Asia. Etymology ''Kiangyousteus'' is named after Jiangyou (formerly ‘Kiangyou’) district of Sichuan Province, the location where the fossils were found in 1953 by Professor S. H. Yoh of Peking University. Phylogeny ''Kiangyousteus'' belongs to the family Dunkleosteidae. The phylogeny of ''Kiangyousteus'' can be shown in the cladogram below: Alternatively, the subsequent 2016 Zhu ''et al.'' study using a larger morphological dataset recovered Panxiosteidae well outside of Dunkleosteoidea, leaving the status of Dunkleosteidae Dunkleosteidae is an extinct family (biology), family of arthrodire placoderms that lived during the Devonian period. The gigantic apex predator ''Dunkleosteus terrelli'' is the best known member of this ...
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