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Fasolasuchus Tenax Skull
''Fasolasuchus'' is an extinct genus of loricatan. Fossils have been found in the Los Colorados Formation of the Ischigualasto-Villa Unión Basin in northwestern Argentina that date back to the Norian stage of the Late Triassic, making it one of the last rauisuchians to have existed before the order (biology), order became extinct at the end of the Triassic. Description It is quite possibly the largest known member of Rauisuchia, with an estimated length of to , even bigger than the Prestosuchidae, prestosuchid ''Saurosuchus'' at in length. This would make ''Fasolasuchus'' the largest terrestrial predator to have ever existed save for large theropods. Like ''Saurosuchus'', it had only a single row of caudal osteoderms, unusual among rauisuchians. It also had a hyposphene-hypantrum articulation that gave the vertebral column extra rigidity. This feature is also seen in several other rauisuchians such as ''Postosuchus'' as well as saurischian dinosaurs. Phylogeny Cladogra ...
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Norian
The Norian is a division of the Triassic Period. It has the rank of an age ( geochronology) or stage (chronostratigraphy). It lasted from ~227 to million years ago. It was preceded by the Carnian and succeeded by the Rhaetian. Stratigraphic definitions The Norian was named after the Noric Alps in Austria. The stage was introduced into scientific literature by Austrian geologist Edmund Mojsisovics von Mojsvar in 1869. The Norian Stage begins at the base of the ammonite biozones of '' Klamathites macrolobatus'' and '' Stikinoceras kerri'', and at the base of the conodont biozones of '' Metapolygnathus communisti'' and '' Metapolygnathus primitius''. A global reference profile for the base (a GSSP) had in 2009 not yet been appointed. The top of the Norian (the base of the Rhaetian) is at the first appearance of ammonite species '' Cochloceras amoenum''. The base of the Rheatian is also close to the first appearance of conodont species '' Misikella spp.'' and '' Epigondolell ...
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Osteoderm
Osteoderms are bony deposits forming scales, plates, or other structures based in the dermis. Osteoderms are found in many groups of extant and extinct reptiles and amphibians, including lizards, crocodilians, frogs, temnospondyls (extinct amphibians), various groups of dinosaurs (most notably ankylosaurs and stegosaurians), phytosaurs, aetosaurs, placodonts, and hupehsuchians (marine reptiles with possible ichthyosaur affinities). Osteoderms are uncommon in mammals, although they have occurred in many xenarthrans ( armadillos and the extinct glyptodonts and mylodontid and scelidotheriid ground sloths). The heavy, bony osteoderms have evolved independently in many different lineages. The armadillo osteoderm is believed to develop in subcutaneous dermal tissues. These varied structures should be thought of as anatomical analogues, not homologues, and do not necessarily indicate monophyly. The structures are however derived from scutes, common to all classes ...
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Paracrocodylomorphs
Paracrocodylomorpha is a clade of pseudosuchian archosaurs. The clade includes the diverse and unusual group Poposauroidea as well as the generally carnivorous and quadrupedal members of Loricata, including modern crocodylians. Paracrocodylomorpha was named by paleontologist J. Michael Parrish in 1993, although the group is now considered to encompass more reptiles than his original definition intended. The most recent definition of Paracrocodylomorpha, as defined by Sterling Nesbitt in 2011, is "the least inclusive clade containing '' Poposaurus'' and ''Crocodylus niloticus'' (the Nile crocodile). Most groups of paracrocodylomorphs became extinct at the end of the Triassic period, with the exception of the crocodylomorphs, from which crocodylians such as crocodiles and alligators evolved in the latter part of the Mesozoic. History and definition Parrish (1993) defined Paracrocodylomorpha as the last common ancestor of " Poposauridae" and Crocodylomorpha and all of its descen ...
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Paleobiology Database
The Paleobiology Database is an online resource for information on the distribution and classification of fossil animals, plants, and microorganisms. History The Paleobiology Database (PBDB) originated in the NCEAS-funded Phanerozoic Marine Paleofaunal Database initiative, which operated from August 1998 through August 2000. From 2000 to 2015, PBDB received funding from the National Science Foundation. PBDB also received support form the Australian Research Council. From 2000 to 2010 it was housed at the National Center for Ecological Analysis and Synthesis, a cross-disciplinary research center within the University of California, Santa Barbara. It is currently housed at University of Wisconsin-Madison and overseen by an international committee of major data contributors. The Paleobiology Database works closely with the Neotoma Paleoecology Database, which has a similar intellectual history, but has focused on the Quaternary (with an emphasis on the late Pleistocene and Holoc ...
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Crocodylomorpha
Crocodylomorpha is a group of pseudosuchian archosaurs that includes the crocodilians and their extinct relatives. They were the only members of Pseudosuchia to survive the end-Triassic extinction. During Mesozoic and early Cenozoic times, crocodylomorphs were far more diverse than they are now. Triassic forms were small, lightly built, active terrestrial animals. The earliest and most primitive crocodylomorphs are represented by " sphenosuchians", a paraphyletic assemblage containing small-bodied forms with elongated limbs that walked upright, which represents the ancestral morphology of Crocodylomorpha. These forms persisted until the end of the Jurassic. During the Jurassic, Crocodylomorphs morphologically diversified into numerous niches, including into the aquatic and marine realms. Evolutionary history When their extinct species and stem group are examined, the crocodylian lineage (clade Pseudosuchia, formerly Crurotarsi) proves to have been a very diverse and adapt ...
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Rauisuchidae
Rauisuchidae is a group of large (up to or more) predatory Triassic archosaurs. There is some disagreement over which genera should be included in Rauisuchidae and which should be in the related Prestosuchidae and Poposauridae, and indeed whether these should even be thought of as separate valid families. Rauisuchids occurred throughout much of the Triassic, and may have first occurred in the Early Triassic if some archosaurian taxa such as ''Scythosuchus'' and ''Tsylmosuchus'' are considered to be within the family. An early cladistic analysis of crocodylotarsan (pseudosuchian) archosaurs included ''Lotosaurus'', ''Fasolasuchus'', ''Rauisuchus'', and "the Kupferzell rauisuchid" (later called ''Batrachotomus'') within Rauisuchidae.Parrish JM. 1993. Phylogeny of the Crocodylotarsi, with reference to archosaurian and crurotarsan monophyly. ''Journal of Vertebrate Paleontology'' 13: 287-308. However, a later study found that ''Batrachotomus'' was a more basal pseudosuchian only sli ...
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Batrachotomus
''Batrachotomus'' is a genus of prehistoric archosaur. Fossils of this animal have been found in southern Germany and dated from the Ladinian, Ladinian stage of the Middle Triassic Geological period, period, around 242 to 237 million years ago. ''Batrachotomus'' was described by palaeontologist David J. Gower 22 years after its discovery. The locality where ''Batrachotomus'' lived was a swampy region and the name comes from the Ancient Greek, Greek ''batrachos/βάτραχος'' (frog) and ''tome/τομή'' (cutting, slicing), which refers to its preying on the large amphibian ''Mastodonsaurus''.Gower (1999), p. 6. In contrast with sprawling reptiles, like crocodiles, this large carnivore was very agile with locomotor superiority due to its erect stance. A remarkable feature seen on its back was a row of paired, flattened bony plates. ''Batrachotomus'' was possibly an early relative of ''Postosuchus'',Gower (1999), p. 1. which lived during the dawn of the dinosaurs. Description ...
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Prestosuchus
''Prestosuchus'' (meaning "Prestes crocodile") is an extinct genus of pseudosuchian in the group Loricata, which also includes ''Saurosuchus'' and ''Postosuchus''. It has historically been referred to as a "rauisuchian", and was the defining member of the family Prestosuchidae, though the validity of both of these groups is questionable: Rauisuchia is now considered paraphyletic and Prestosuchidae is polyphyletic in its widest form. History of study ''Prestosuchus chiniquensis'' was first discovered in the Santa Maria Formation at the Paleontological Site Chiniquá, near the city of São Pedro do Sul in 1928 or 1929, by the German paleontologist Friedrich von Huene on a trip to Brazil. Von Huene named the genus in 1938 in honor of Vicentino Prestes de Almeida. This site is located in the geopark of Paleorrota. Munich specimens The first two specimens of ''Prestosuchus'' to be described were found at the Weg Sanga site near the town of Sao Pedro do Sul in Rio Grande do S ...
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Loricata
Loricata is a clade of archosaur reptiles that includes crocodilians and some of their Triassic relatives, such as ''Postosuchus'' and ''Prestosuchus''. More specifically, Loricata includes Crocodylomorpha (the persistent archosaur subset which crocodilians belong to) and most " rauisuchians", a paraphyletic grade of large terrestrial pseudosuchians which were alive in the Triassic period and ancestral to crocodylomorphs. Loricata is one branch of the group Paracrocodylomorpha; the other branch is the clade Poposauroidea, an unusual collection of strange "rauisuchians" including bipedal, herbivorous, and sail-backed forms. The vast majority of typical "rauisuchians", which were usually quadrupedal predators, occupy basal (early-branching) rungs of Loricata leading up to crocodylomorphs. History and Classification Loricata was initially named in a completely different context by German naturalist Blasius Merrem in his 1820 ''Versuch eines Systems der Amphibien''. Merrem consi ...
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Poposauroidea
Poposauroidea is a clade of advanced pseudosuchians (archosaurs closer to crocodilians than to dinosaurs). It includes poposaurids, shuvosaurids, ctenosauriscids, and other unusual pseudosuchians such as ''Qianosuchus'' and '' Lotosaurus''. However, it excludes most large predatory quadrupedal " rauisuchians" such as rauisuchids and " prestosuchids". Those reptiles are now allied with crocodylomorphs (crocodile ancestors) in a clade known as Loricata, which is the sister taxon to the poposauroids in the clade Paracrocodylomorpha. Although it was first formally defined in 2007, the name "Poposauroidea" has been used for many years. The group has been referred to as Poposauridae by some authors, although this name is often used more narrowly to refer to the family that includes ''Poposaurus'' and its close relatives. Poposauroids went extinct at the end of the Triassic period along with other non-crocodylomorph pseudosuchians. However, they were among the most diverse and lon ...
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Paracrocodylomorpha
Paracrocodylomorpha is a clade of pseudosuchian archosaurs. The clade includes the diverse and unusual group Poposauroidea as well as the generally carnivorous and quadrupedal members of Loricata, including modern crocodylians. Paracrocodylomorpha was named by paleontologist J. Michael Parrish in 1993, although the group is now considered to encompass more reptiles than his original definition intended. The most recent definition of Paracrocodylomorpha, as defined by Sterling Nesbitt in 2011, is "the least inclusive clade containing '' Poposaurus'' and ''Crocodylus niloticus'' (the Nile crocodile). Most groups of paracrocodylomorphs became extinct at the end of the Triassic period, with the exception of the crocodylomorphs, from which crocodylians such as crocodiles and alligators evolved in the latter part of the Mesozoic. History and definition Parrish (1993) defined Paracrocodylomorpha as the last common ancestor of " Poposauridae" and Crocodylomorpha and all of its des ...
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Cladogram
A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to descendants, nor does it show how much they have changed, so many differing evolutionary trees can be consistent with the same cladogram. A cladogram uses lines that branch off in different directions ending at a clade, a group of organisms with a last common ancestor. There are many shapes of cladograms but they all have lines that branch off from other lines. The lines can be traced back to where they branch off. These branching off points represent a hypothetical ancestor (not an actual entity) which can be inferred to exhibit the traits shared among the terminal taxa above it. This hypothetical ancestor might then provide clues about the order of evolution of various features, adaptation, and other evolutionary narratives about an ...
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