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Evolution Of Mollusca
The evolution of the molluscs is the way in which the Mollusca, one of the largest groups of invertebrate animals, evolved. This phylum includes gastropods, bivalves, scaphopods, cephalopods, and several other groups. The fossil record of mollusks is relatively complete, and they are well represented in most fossil-bearing marine strata. Very early organisms which have dubiously been compared to molluscs include ''Kimberella'' and '' Odontogriphus''. Fossil record Good evidence exists for the appearance of gastropods, cephalopods and bivalves in the Cambrian period . However, the evolutionary history both of the emergence of molluscs from the ancestral group Lophotrochozoa, and of their diversification into the well-known living and fossil forms, is still vigorously debated. Debate occurs about whether some Ediacaran and Early Cambrian fossils really are molluscs. ''Kimberella'', from about , has been described by some paleontologists as "mollusc-like", but others are unw ...
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Cambrian
The Cambrian Period ( ; sometimes symbolized Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million years ago (mya) to the beginning of the Ordovician Period mya. Its subdivisions, and its base, are somewhat in flux. The period was established as "Cambrian series" by Adam Sedgwick, who named it after Cambria, the Latin name for 'Cymru' (Wales), where Britain's Cambrian rocks are best exposed. Sedgwick identified the layer as part of his task, along with Roderick Murchison, to subdivide the large "Transition Series", although the two geologists disagreed for a while on the appropriate categorization. The Cambrian is unique in its unusually high proportion of sedimentary deposits, sites of exceptional preservation where "soft" parts of organisms are preserved as well as their more resistant shells. As a result, our understanding of the Cambria ...
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Siphuncle
The siphuncle is a strand of tissue passing longitudinally through the shell of a cephalopod mollusk. Only cephalopods with chambered shells have siphuncles, such as the extinct ammonites and belemnites, and the living nautiluses, cuttlefish, and ''Spirula''. In the case of the cuttlefish, the siphuncle is indistinct and connects all the small chambers of that animal's highly modified shell; in the other cephalopods it is thread-like and passes through small openings in the septa (walls) dividing the camerae (chambers). Some older studies have used the term siphon for the siphuncle, though this naming convention is uncommon in modern studies to prevent confusion with a mollusc organ of the same name. Function The siphuncle is used primarily in emptying water from new chambers as the shell grows. To perform this task, the cephalopod increases the saltiness of the blood in the siphuncle, and the water moves from the more dilute chamber into the blood through osmosis. At the sa ...
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Torsion (gastropod)
Torsion is a gastropod synapomorphy which occurs in all gastropods during larval development. Torsion is the rotation of the visceral mass, mantle, and shell 180˚ with respect to the head and foot of the gastropod. This rotation brings the mantle cavity and the anus to an anterior position above the head. In some groups of gastropods (Opisthobranchia) there is a degree of secondary detorsion or rotation towards the original position; this may be only partial detorsion or full detorsion. The torsion or twisting of the visceral mass of larval gastropods is not the same thing as the spiral coiling of the shell, which is also present in many shelled gastropods. Development There are two different developmental stages which cause torsion. The first stage is caused by the development of the asymmetrical velar/foot muscle which has one end attached to the left side of the shell and the other end has fibres attached to the left side of the foot and head. At a certain point in larva ...
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Limpet
Limpets are a group of aquatic snails that exhibit a conical gastropod shell, shell shape (patelliform) and a strong, muscular foot. Limpets are members of the class Gastropoda, but are polyphyletic, meaning the various groups called "limpets" descended independently from different ancestral gastropods. This general category of conical shell is known as "patelliform" (dish-shaped). All members of the large and ancient marine clade Patellogastropoda are limpets. Within that clade, the members of the Patellidae family in particular are often referred to as "true limpets". Other groups, not in the same family, are also called limpets of one type or another, due to the similarity of their shells' shape. Examples include the Fissurellidae ("keyhole limpet") family, which is part of the Vetigastropoda clade (many other members of the Vetigastropoda do not have the Morphology (biology), morphology of limpets) and the Siphonariidae ("false limpets"), which use a siphon to pump water o ...
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Trilobite
Trilobites (; meaning "three lobes") are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest-known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the Atdabanian stage of the Early Cambrian period () and they flourished throughout the lower Paleozoic before slipping into a long decline, when, during the Devonian, all trilobite orders except the Proetida died out. The last extant trilobites finally disappeared in the mass extinction at the end of the Permian about 252 million years ago. Trilobites were among the most successful of all early animals, existing in oceans for almost 270 million years, with over 22,000 species having been described. By the time trilobites first appeared in the fossil record, they were already highly diversified and geographically dispersed. Because trilobites had wide diversity and an easily fossilized exoskeleton, they left an extensive fossil record. Th ...
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China
China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's List of countries and dependencies by population, most populous country, with a Population of China, population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and Borders of China, borders fourteen countries by land, the List of countries and territories by land borders, most of any country in the world, tied with Russia. Covering an area of approximately , it is the world's third List of countries and dependencies by area, largest country by total land area. The country consists of 22 provinces of China, provinces, five autonomous regions of China, autonomous regions, four direct-administered municipalities of China, municipalities, and two special administrative regions of China, Special Administrative Regions (Hong Kong and Macau). The national capital is Beijing, and the List of cities in China by population, most populous cit ...
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Siberia
Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part of Russia since the latter half of the 16th century, after the Russians conquered lands east of the Ural Mountains. Siberia is vast and sparsely populated, covering an area of over , but home to merely one-fifth of Russia's population. Novosibirsk, Krasnoyarsk and Omsk are the largest cities in the region. Because Siberia is a geographic and historic region and not a political entity, there is no single precise definition of its territorial borders. Traditionally, Siberia extends eastwards from the Ural Mountains to the Pacific Ocean, and includes most of the drainage basin of the Arctic Ocean. The river Yenisey divides Siberia into two parts, Western and Eastern. Siberia stretches southwards from the Arctic Ocean to the hills of nort ...
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Microfossil
A microfossil is a fossil that is generally between 0.001 mm and 1 mm in size, the visual study of which requires the use of light or electron microscopy. A fossil which can be studied with the naked eye or low-powered magnification, such as a hand lens, is referred to as a macrofossil. Microfossils are a common feature of the geological record, from the Precambrian to the Holocene. They are most common in deposits of marine environments, but also occur in brackish water, fresh water and terrestrial sedimentary deposits. While every kingdom of life is represented in the microfossil record, the most abundant forms are protist skeletons or microbial cysts from the Chrysophyta, Pyrrhophyta, Sarcodina, acritarchs and chitinozoans, together with pollen and spores from the vascular plants. Overview A microfossil is a descriptive term applied to fossilized plants and animals whose size is just at or below the level at which the fossil can be analyzed by the naked eye. A ...
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Polychaete
Polychaeta () is a paraphyletic class of generally marine annelid worms, commonly called bristle worms or polychaetes (). Each body segment has a pair of fleshy protrusions called parapodia that bear many bristles, called chaetae, which are made of chitin. More than 10,000 species are described in this class. Common representatives include the lugworm (''Arenicola marina'') and the sandworm or clam worm ''Alitta''. Polychaetes as a class are robust and widespread, with species that live in the coldest ocean temperatures of the abyssal plain, to forms which tolerate the extremely high temperatures near hydrothermal vents. Polychaetes occur throughout the Earth's oceans at all depths, from forms that live as plankton near the surface, to a 2- to 3-cm specimen (still unclassified) observed by the robot ocean probe ''Nereus'' at the bottom of the Challenger Deep, the deepest known spot in the Earth's oceans. Only 168 species (less than 2% of all polychaetes) are known from ...
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Radula
The radula (, ; plural radulae or radulas) is an anatomical structure used by molluscs for feeding, sometimes compared to a tongue. It is a minutely toothed, chitinous ribbon, which is typically used for scraping or cutting food before the food enters the esophagus. The radula is unique to the molluscs, and is found in every class of mollusc except the bivalves, which instead use cilia, waving filaments that bring minute organisms to the mouth. Within the gastropods, the radula is used in feeding by both herbivorous and carnivorous snails and slugs. The arrangement of teeth ( denticles) on the radular ribbon varies considerably from one group to another. In most of the more ancient lineages of gastropods, the radula is used to graze, by scraping diatoms and other microscopic algae off rock surfaces and other substrates. Predatory marine snails such as the Naticidae use the radula plus an acidic secretion to bore through the shell of other molluscs. Other predatory marine sn ...
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Wiwaxia
''Wiwaxia'' is a genus of soft-bodied animals that were covered in carbonaceous scales and spines that protected it from predators. ''Wiwaxia'' fossils – mainly isolated scales, but sometimes complete, articulated fossils – are known from early Cambrian and middle Cambrian fossil deposits across the globe. The living animal would have measured up to 5 cm (2 inch) when fully grown, although a range of juvenile specimens are known, the smallest being long. ''Wiwaxia's'' affinity has been a matter of debate: Researchers were long split between two possibilities. On the one hand, its rows of scales looked superficially similar to certain scale worms (annelids); conversely, its mouthparts and general morphology suggested a relationship to the shell-less molluscs. More recently, evidence for a molluscan affinity has been accumulating, based on new details of ''Wiwaxias mouthparts, scales, and growth history. The proposed clade Halwaxiida contains ''Wiwaxia'' as well a ...
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