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Eucacopinae
Eucacopinae is an extinct clade (evolutionary grouping) of dissorophid temnospondyls. Eucacopines differ from the other main group of dissorophids, the Dissorophinae, in having more lightly built skeletons and more knobby skulls. The subfamily was originally named Cacopinae, but since the name was already established for a group of living microhylid frogs in 1931, the name was changed to Eucacopinae in 2013. Eucacopinae is a stem-based taxon defined as the most inclusive clade containing the species '' Cacops apsidephorus'' but not ''Dissorophus multicinctus'', which belongs to Dissorophinae. According to the most recent phylogenetic analyses of Dissorophidae, Eucacopinae includes the basal ("primitive") species '' Conjunctio multidens'' and '' Scapanops neglecta'' from the southwestern United States and a more derived ("advanced") group including several species of ''Cacops'' (also from the southwestern United States) and the Russian genera '' Kamacops'' and '' Zygosaurus''. D ...
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Scapanops Neglecta
''Scapanops'' is an extinct genus of dissorophid temnospondyl amphibian known from the Early Permian Nocona Formation of north-central Texas, United States. It contains only the type species ''Scapanops neglecta'', which was named by Rainer R. Schoch and Hans-Dieter Sues in 2013. ''Scapanops'' differs from other dissorophids in having a very small skull table, which means that its eye sockets are unusually close to the back of the skull. The eye sockets are also very large and spaced far apart. ''Scapanops'' was probably small-bodied (around long) with a proportionally large head and short trunk and tail. Like other dissorophids, it probably spent most of its life on land. Description The only known skull of ''Scapanops'' is long. ''Scapanops'' is distinguished from other dissorophids by five unique features or autapomorphies: a large spacing between the orbits or eye sockets, giving the skull an oval-shaped outline; a very small skull table behind the orbits; a long snout t ...
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Dissorophid
Dissorophidae is an extinct family of medium-sized, temnospondyl amphibians that flourished during the late Carboniferous and early Permian periods. The clade is known almost exclusively from North America. History of study Dissorophidae is a diverse clade that was named in 1902 by George A. Boulenger. Junior synonyms include Otocoelidae, Stegopidae, and Aspidosauridae. Early in the study of dissorophoids when the relationships of different taxa were not well-resolved and most taxa had not been described, Dissorophidae sometimes came to include taxa that are now not regarded as dissorophids and may have excluded earlier described taxa that are now regarded as dissorophids. Amphibamiforms were widely regarded as small-bodied dissorophids, and at one point, Dissorophidae was also suggested to also include Trematopidae. 19th century In 1895, American paleontologist Edward Drinker Cope named '' Dissorophus'' from the early Permian of Texas. This was the first dissorophid to be ...
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Dissorophids
Dissorophidae is an extinct family of medium-sized, temnospondyl amphibians that flourished during the late Carboniferous and early Permian periods. The clade is known almost exclusively from North America. History of study Dissorophidae is a diverse clade that was named in 1902 by George A. Boulenger. Junior synonyms include Otocoelidae, Stegopidae, and Aspidosauridae. Early in the study of dissorophoids when the relationships of different taxa were not well-resolved and most taxa had not been described, Dissorophidae sometimes came to include taxa that are now not regarded as dissorophids and may have excluded earlier described taxa that are now regarded as dissorophids. Amphibamiforms were widely regarded as small-bodied dissorophids, and at one point, Dissorophidae was also suggested to also include Trematopidae. 19th century In 1895, American paleontologist Edward Drinker Cope named ''Dissorophus'' from the early Permian of Texas. This was the first dissorophid to be de ...
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Temnospondyl
Temnospondyli (from Greek τέμνειν, ''temnein'' 'to cut' and σπόνδυλος, ''spondylos'' 'vertebra') is a diverse order of small to giant tetrapods—often considered primitive amphibians—that flourished worldwide during the Carboniferous, Permian, and Triassic periods. A few species continued into the Jurassic and Cretaceous periods. Fossils have been found on every continent. During about 210 million years of evolutionary history, they adapted to a wide range of habitats, including freshwater, terrestrial, and even coastal marine environments. Their life history is well understood, with fossils known from the larval stage, metamorphosis, and maturity. Most temnospondyls were semiaquatic, although some were almost fully terrestrial, returning to the water only to breed. These temnospondyls were some of the first vertebrates fully adapted to life on land. Although temnospondyls are considered amphibians, many had characteristics, such as scales and armour-like bo ...
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Conjunctio Multidens
''Conjunctio'' is an extinct genus of dissorophid temnospondyl amphibian from the early Permian of New Mexico. The type species, ''Conjunctio multidens'', was named by paleontologist Robert L. Carroll in 1964. History of study The holotype specimen was found in 1911 in the Lower Permian Abo Formation in New Mexico ) , population_demonym = New Mexican ( es, Neomexicano, Neomejicano, Nuevo Mexicano) , seat = Santa Fe, New Mexico, Santa Fe , LargestCity = Albuquerque, New Mexico, Albuquerque , LargestMetro = Albuquerque metropolitan area, Tiguex , Offi .... It consists of the skull and postcranial material including femora, humeri, scapulae, pelvis, a section of the vertebral column, and osteoderms. It was originally described by Case et al. (1913) as a referred specimen of ''Aspidosaurus'' (sometimes "''Broiliellus''") ''novomexicanus'' but was subsequently determined to be a distinct species by Carroll (1964). Carroll also identified a third specimen of ''Conjunctio'', ...
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Late Carboniferous
Late may refer to: * LATE, an acronym which could stand for: ** Limbic-predominant age-related TDP-43 encephalopathy, a proposed form of dementia ** Local-authority trading enterprise, a New Zealand business law ** Local average treatment effect, a concept in econometrics Music * ''Late'' (album), a 2000 album by The 77s * Late!, a pseudonym used by Dave Grohl on his '' Pocketwatch'' album * Late (rapper), an underground rapper from Wolverhampton * "Late" (song), a song by Blue Angel * "Late", a song by Kanye West from '' Late Registration'' Other * Late (Tonga), an uninhabited volcanic island southwest of Vavau in the kingdom of Tonga * "Late" (''The Handmaid's Tale''), a television episode * LaTe, Oy Laivateollisuus Ab, a defunct shipbuilding company * Late may refer to a person who is Dead See also * * * '' Lates'', a genus of fish in the lates perch family * Later (other) * Tardiness * Tardiness (scheduling) {{disambig ...
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Basal (phylogenetics)
In phylogenetics, basal is the direction of the ''base'' (or root) of a rooted phylogenetic tree or cladogram. The term may be more strictly applied only to nodes adjacent to the root, or more loosely applied to nodes regarded as being close to the root. Note that extant taxa that lie on branches connecting directly to the root are not more closely related to the root than any other extant taxa. While there must always be two or more equally "basal" clades sprouting from the root of every cladogram, those clades may differ widely in taxonomic rank, species diversity, or both. If ''C'' is a basal clade within ''D'' that has the lowest rank of all basal clades within ''D'', ''C'' may be described as ''the'' basal taxon of that rank within ''D''. The concept of a 'key innovation' implies some degree of correlation between evolutionary innovation and diversification. However, such a correlation does not make a given case predicable, so ancestral characters should not be imputed ...
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Most Recent Common Ancestor
In biology and genetic genealogy, the most recent common ancestor (MRCA), also known as the last common ancestor (LCA) or concestor, of a set of organisms is the most recent individual from which all the organisms of the set are descended. The term is also used in reference to the ancestry of groups of genes ( haplotypes) rather than organisms. The MRCA of a set of individuals can sometimes be determined by referring to an established pedigree. However, in general, it is impossible to identify the exact MRCA of a large set of individuals, but an estimate of the time at which the MRCA lived can often be given. Such ''time to most recent common ancestor'' (''TMRCA'') estimates can be given based on DNA test results and established mutation rates as practiced in genetic genealogy, or by reference to a non-genetic, mathematical model or computer simulation. In organisms using sexual reproduction, the ''matrilineal MRCA'' and ''patrilineal MRCA'' are the MRCAs of a given populati ...
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Osteoderm
Osteoderms are bony deposits forming scales, plates, or other structures based in the dermis. Osteoderms are found in many groups of extant and extinct reptiles and amphibians, including lizards, crocodilians, frogs, temnospondyls (extinct amphibians), various groups of dinosaurs (most notably ankylosaurs and stegosaurians), phytosaurs, aetosaurs, placodonts, and hupehsuchians (marine reptiles with possible ichthyosaur affinities). Osteoderms are uncommon in mammals, although they have occurred in many xenarthrans ( armadillos and the extinct glyptodonts and mylodontid and scelidotheriid ground sloths). The heavy, bony osteoderms have evolved independently in many different lineages. The armadillo osteoderm is believed to develop in subcutaneous dermal tissues. These varied structures should be thought of as anatomical analogues, not homologues, and do not necessarily indicate monophyly. The structures are however derived from scutes, common to all classes ...
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Zygosaurus
''Zygosaurus'' is an extinct genus of dissorophid temnospondyl from the Middle-Late Permian of Russia. It was described in 1848 by Eduard Eichwald, making it the first dissorophid to be described and is known from a single species, ''Zygosaurus lucius''. The location of the holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of seve ..., and only known specimen, is unknown, and although casts are reposited in several institutions, little is known about this taxon beyond qualitative aspects of the skull (e.g., preorbital length twice as long as postorbital length; skull width greatest at mid-length of orbits). The skull was estimated to be around 20 cm in length, making it one of the largest dissorophids, being only slightly smaller than '' Kamacops''. References Dissorophi ...
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Kamacops
''Kamacops'' is a genus of dissorophid temnospondyls known from the Middle to Late Permian of Russia that was described by Yuri Gubin in 1980. It is known from a single species, ''Kamacops acervalis,'' material of which is currently reposited in the Paleontological Institute of the Russian Academy of Sciences. ''Kamacops'' is one of the youngest dissorophids, along with '' Iratusaurus'' and ''Zygosaurus'' from Russia and '' Anakamacops'' from China and was one of the largest known dissorophids, with an estimated skull length of 24–30 cm. It is typically recovered as being most closely related to ''Zygosaurus'' and to the North American ''Cacops ''Cacops'' ("ugly look" for its strange appearance), is a genus of dissorophid temnospondyls from the Kungurian stage of the early Permian of the United States. ''Cacops'' is one of the few olsoniforms (dissorophids and the larger trematopids ...''''.'' A detailed study of the braincase region was performed by Schoch (1999) ...
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Synapomorphy
In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to have evolved in their most recent common ancestor. ) In cladistics, synapomorphy implies homology. Examples of apomorphy are the presence of erect gait, fur, the evolution of three middle ear bones, and mammary glands in mammals but not in other vertebrate animals such as amphibians or reptiles, which have retained their ancestral traits of a sprawling gait and lack of fur. Thus, these derived traits are also synapomorphies of mammals in general as they are not shared by other vertebrate animals. Etymology The word —coined by German entomologist Willi Hennig—is derived from the Ancient Greek words (''sún''), meaning "with, together"; (''apó''), meaning "away from"; and (''morphḗ''), meaning "shape, form". Clade ana ...
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