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Euarchontoglires
Euarchontoglires (from: '' Euarchonta'' ("true rulers") + '' Glires'' ("dormice")), synonymous with Supraprimates, is a clade and a superorder of placental mammals, the living members of which belong to one of the five following groups: rodents, lagomorphs, treeshrews, primates, and colugos. Evolutionary affinities within mammals The Euarchontoglires clade is based on DNA sequence analyses and retrotransposon markers that combine the clades Glires (Rodentia + Lagomorpha) and Euarchonta (Scandentia + Primates + Dermoptera). It is usually discussed without a taxonomic rank but has been called a cohort, magnorder, or superorder. Relations among the four cohorts (Euarchontoglires, Xenarthra, Laurasiatheria, Afrotheria) and the identity of the placental root remain controversial. So far, few, if any, distinctive anatomical features have been recognized that support Euarchontoglires; nor does any strong evidence from anatomy support alternative hypotheses. Although both Eu ...
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Euarchonta
The Euarchonta are a proposed grandorder of mammals: the order Scandentia (treeshrews), and its sister Primatomorpha mirorder, containing the Dermoptera (colugos) and the primates (Plesiadapiformes and descendants). The term "Euarchonta" (meaning "true rulers") appeared in 1999, when molecular evidence suggested that the morphology-based Archonta should be trimmed down to exclude Chiroptera. Further DNA sequence analyses supported the Euarchonta hypothesis. Despite multiple papers pointing out that some mitochondrial sequences showed unusual properties (particularly murid rodents and hedgehogs) and were likely distorting the overall tree, and despite earlier studies showing near total congruence of mtDNA-based and nuclear-based trees when such sequences were excluded, some authors continued to produce misleading trees. A study investigating retrotransposon presence/absence data has claimed strong support for Euarchonta. Some interpretations of the molecular data link Primates ...
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Treeshrew
The treeshrews (also called tree shrews or banxrings) are small mammals native to the tropical forests of South and Southeast Asia. They make up the entire order Scandentia (from Latin ''scandere'', "to climb"), which split into two families: the Tupaiidae (19 species, "ordinary" treeshrews), and the Ptilocercidae (one species, the pen-tailed treeshrew). Though called 'treeshrews', and despite having previously been classified in Insectivora, they are not true shrews, and not all species live in trees. They are omnivores; among other things, treeshrews eat fruit. As fellow members of Euarchonta, treeshrews are closely related to primates, and have been used as an alternative to primates in experimental studies of myopia, psychosocial stress, and hepatitis. Description Treeshrews are slender animals with long tails and soft, greyish to reddish-brown fur. The terrestrial species tend to be larger than the arboreal forms, and to have larger claws, which they use for digging up ...
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Glires
Glires (, Latin ''glīrēs'' 'dormice') is a clade (sometimes ranked as a grandorder) consisting of rodents and lagomorphs ( rabbits, hares, and pikas). The hypothesis that these form a monophyletic group has been long debated based on morphological evidence. Two morphological studies, published in 2001 and 2003, strongly support the monophyly of Glires. In particular, the 2003 study reported the discovery of fossil material of basal members of Glires, particularly the genera '' Mimotona'', '' Gomphos'', '' Heomys'', '' Matutinia'', '' Rhombomylus'', and '' Sinomylus''. Their description, in 2005, helped to bridge the gap between more typical rodents and lagomorphs. Data published in 2001, based on nuclear DNA, supported Glires as a sister of Euarchonta to form Euarchontoglires, but some genetic data from both nuclear and mitochondrial DNA Mitochondrial DNA (mtDNA and mDNA) is the DNA located in the mitochondrion, mitochondria organelles in a eukaryotic cell that c ...
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Primate
Primates is an order (biology), order of mammals, which is further divided into the Strepsirrhini, strepsirrhines, which include lemurs, galagos, and Lorisidae, lorisids; and the Haplorhini, haplorhines, which include Tarsiiformes, tarsiers and simians (monkeys and apes). Primates arose 74–63 million years ago first from small terrestrial animal, terrestrial mammals, which adapted for life in tropical forests: many primate characteristics represent adaptations to the challenging environment among Canopy (biology), tree tops, including large brain sizes, binocular vision, color vision, Animal communication, vocalizations, shoulder girdles allowing a large degree of movement in the upper limbs, and opposable thumbs (in most but not all) that enable better grasping and dexterity. Primates range in size from Madame Berthe's mouse lemur, which weighs , to the eastern gorilla, weighing over . There are 376–524 species of living primates, depending on which classification is ...
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Colugo
Colugos (), flying lemurs, or cobegos (), are arboreal gliding euarchontogliran mammals that are native to Southeast Asia. Their closest evolutionary relatives are primates. There are just two living species of colugos: the Sunda flying lemur (''Galeopterus variegatus'') and the Philippine flying lemur (''Cynocephalus volans''). These two species make up the entire family Cynocephalidae () and order Dermoptera, from Ancient Greek δέρμα (''dérma'') "skin" and πτερόν (''pterón'') "wing". Characteristics Colugos are nocturnal, tree-dwelling mammals. Appearance and anatomy They reach lengths of and weigh . They have long, slender front and rear limbs, a medium-length tail, and a relatively light build. The head is small, with large, front-focused eyes for excellent binocular vision, and small rounded ears. The incisor teeth of colugos are highly distinctive; they are comb-like in shape with up to 20 tines on each tooth. The incisors are analogous in app ...
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Apatemyidae
Apatemyidae is an extinct family of placental mammals that took part in the first placental evolutionary radiation together with other early mammals, such as the leptictids. Their relationships to other mammal groups are controversial; a 2010 study found them to be basal members of Euarchontoglires. Common in North America during the Paleocene, they are also represented in Europe by the genus '' Jepsenella''. Apatemyids in life Like most Paleocene mammals, the apatemyids were small and presumably insectivorous. Size ranged from that of a dormouse to a large rat. The toes were slender and well clawed, and the family were probably mainly arboreal. The skull was fairly massive compared to the otherwise slender skeleton, and the front teeth were long and hooked, resembling those of the modern aye-aye The aye-aye (''Daubentonia madagascariensis'') is a long-fingered lemur, a Strepsirrhini, strepsirrhine primate native to Madagascar with rodent-like teeth that perpetually grow a ...
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Laurasiatheria
Laurasiatheria (; "Laurasian beasts") is a superorder of Placentalia, placental mammals that groups together true insectivores (eulipotyphlans), bats (chiropterans), carnivorans, pangolins (Pholidota, pholidotes), even-toed ungulates (Artiodactyla, artiodactyls), odd-toed ungulates (Perissodactyla, perissodactyls), and all their extinct relatives (Pan-Euungulata, pan-euungulates). From systematics and phylogenetic perspectives, it is subdivided into order Eulipotyphla and clade Scrotifera. It is a sister group to Euarchontoglires with which it forms the magnorder Boreoeutheria. Laurasiatheria was discovered on the basis of the Molecular phylogenetics, similar gene sequences shared by the mammals belonging to it; no Anatomy, anatomical features have yet been found that unite the group, although a few have been suggested such as a small coracoid process, a simplified hindgut (reversed in artiodactyls), high intelligence, lack of grasping hands (though mimicry of grasping is observed ...
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Xenarthra
Xenarthra (; from Ancient Greek ξένος, xénos, "foreign, alien" + ἄρθρον, árthron, "joint") is a superorder and major clade of placental mammals native to the Americas. There are 31 living species: the anteaters, tree sloths, and armadillos. Extinct xenarthrans include the glyptodonts, pampatheres and ground sloths. Xenarthrans originated in South America during the late Paleocene about 60 million years ago. They evolved and diversified extensively in South America during the continent's long period of isolation in the early to mid Cenozoic Era. They spread to the Antilles by the early Miocene and, starting about 3 million years ago, spread to Central and North America as part of the Great American Interchange. Nearly all of the formerly abundant megafaunal xenarthrans became extinct at the end of the Pleistocene. Characteristics Xenarthrans share several characteristics that are not present in other placental mammals, which suggest that xenarthrans desce ...
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Anagaloidea
Anagaloidea is a former order of extinct placental mammals that first appeared during the Paleocene epoch. Taxonomy According to the traditional (morphological) view, Anagaloidea is part of the superorder Anagalida, along with the elephant shrews, rodents and lagomorphs. However, the Anagalida are considered to be polyphyletic. Genetic studies have shown that the elephant shrews are actually part of a different macro-group of mammals called the Afrotheria, while the position of several extinct families of Anagalida is uncertain. The Zalambdalestidae are almost certainly unrelated to any of these groups; they probably represent more basal Eutherians and might not even be true Eutherians at all. The Anagalidae and the Pseudictopidae probably represent a genuine clade. This clade is known as the Anagaloidea, which seems to be related to the rodents and lagomorphs after all. Together they form the clade Glires, often grouped with the Euarchonta to form the superorder Euarchontogl ...
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Arctostylopidae
Arctostylopidae is an extinct family of placental mammals from the Late Palaeocene of Eastern Asia and North America. All arctostylopid specimens in North America have been referred to the genus '' Arctostylops''. They are animals of uncertain affinities to other groups and it was believed that they may be related to 'ungulate Ungulates ( ) are members of the diverse clade Euungulata ("true ungulates"), which primarily consists of large mammals with Hoof, hooves. Once part of the clade "Ungulata" along with the clade Paenungulata, "Ungulata" has since been determined ...s'. Originally they were considered to be northern relatives of South American notoungulates, specifically Notostylopidae. Recently, other palaeontologists have suggested that they might be descendants of Asian gliriforms, and therefore related to rabbits and rodents. This relationship is based upon similarities in the shape of their tarsal (ankle) bones. In particular, arctostylopid tarsals bear a strong r ...
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Primatomorpha
Primatomorpha is a proposed mirorder of mammals containing the orders Dermoptera (or colugos) and Primates. Primatomorpha is sister to Scandentia, together forming the Euarchonta. The term "Primatomorpha" first appeared in the general scientific literature in 1991 (K.C. Beard) and 1992 (Kalandadze, Rautian). Major DNA sequence analyses of predominantly nuclear sequences (Murphy et al., 2001) support the Euarchonta hypothesis, while a major study investigating mitochondrial sequences supports a different tree topology (Arnason et al., 2002). A study investigating retrotransposon presence/absence data has claimed strong support for Euarchonta (Kriegs et al., 2007). Some interpretations of the molecular data link Primates and Dermoptera in a clade In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a ...
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Placental Mammal
Placental mammals ( infraclass Placentalia ) are one of the three extant subdivisions of the class Mammalia, the other two being Monotremata and Marsupialia. Placentalia contains the vast majority of extant mammals, which are partly distinguished from monotremes and marsupials in that the fetus is carried in the uterus of its mother to a relatively late stage of development. The name is something of a misnomer, considering that marsupials also nourish their fetuses via a placenta, though for a relatively briefer period, giving birth to less-developed young, which are then nurtured for a period inside the mother's pouch. Placentalia represents the only living group within Eutheria, which contains all mammals that are more closely related to placentals than they are to marsupials. Anatomical features Placental mammals are anatomically distinguished from other mammals by: * a sufficiently wide opening at the bottom of the pelvis to allow the birth of a large baby relative to the ...
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