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Estemmenosuchidae
Estemmenosuchidae is an extinct family of large, very early herbivorous therapsids that flourished during the Guadalupian period. They are distinguished by horn-like structures, probably for display or agonistic behavior. Apart from the best known genus, '' Estemmenosuchus'', the group is poorly known. To date, their fossils are known only from the Perm region of Russia (a region referred to by Russian paleontologists as the Cis-Urals). Description Estemmenosuchids are among the most distinctive of the Permian tetrapods. The high and massive skull is equipped with a number of horns projecting both upwards and outwards, which were probably used for intra-specific display. The incisors and canine teeth are large, but those at the side are reduced, with a serrated apex, and may have helped to break up plant material, although they were too small to be of much use. The body is large and bulky, indicating a large digestive tract for digesting volumes of plant food. The skull supe ...
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Dinocephalia
Dinocephalians (terrible heads) are a clade of large-bodied early therapsids that flourished in the Early and Middle Permian between 279.5 and 260 million years ago (Ma), but became extinct during the Capitanian mass extinction event. Dinocephalians included herbivorous, carnivorous, and omnivorous forms. Many species had thickened skulls with many knobs and bony projections. Dinocephalians were the first non-mammalian therapsids to be scientifically described and their fossils are known from Russia, China, Brazil, South Africa, Zimbabwe, and Tanzania. Description Apart from the biarmosuchians, the dinocephalians are the least advanced therapsids, although still uniquely specialised in their own way. They retain a number of primitive characteristics (e.g. no secondary palate, small dentary) shared with their pelycosaur ancestors, although they are also more advanced in possessing therapsid adaptations like the expansion of the ilium and more erect limbs. They ...
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Anteosaur
Anteosaurs are a group of large, primitive carnivorous dinocephalian therapsids with large canines and incisors and short limbs, that are known from the Middle Permian of South Africa, Russia, China, and Brazil. Some grew very large, with skulls long, and were the largest predators of their time. They died out at the end of the Middle Permian, possibly as a result of the extinction of the herbivorous Tapinocephalia on which they may have fed. Description The Anteosauria are distinguished from the Tapinocephalia by several features, such as very large canines, cheek teeth with bulbous crowns, and an upturning of the premaxilla, so that the front of mouth curves strongly upwards. There is a tendency especially in more advanced forms such as ''Anteosaurus'' towards thickening of the bones of the top of the skull, indicating head-butting behavior. There is a large canal for the pineal organ (third eye); probably tied in with the animal's diurnal and seasonal cycles. The shoulde ...
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Estemmenosuchus
''Estemmenosuchus'' (meaning "crowned crocodile" in Greek language, Greek) is an extinct genus of large, early omnivore, omnivorous therapsid. It is believed and interpreted to have lived during the middle part of the Middle Permian around 267 million years ago. The two species, ''E. uralensis'' and ''E. mirabilis'', are characterised by distinctive horn-like structures, which were probably used for display (zoology), intra-specific display. Both species of ''Estemmenosuchus'' are from the Perm, Russia, Perm (or Cis-Urals) region of Russia. Two other estemmenosuchids, ''Anoplosuchus'' and ''Zopherosuchus'', are now considered females of the species ''E. uralensis''. There were many complete and incomplete skeletons found together. Description ''Estemmenosuchus'' could reach a body length of more than . Its skull was long and massive, up to in length, and possessed several sets of large horns, somewhat similar to the antlers of a moose, growing upward and outward from the sides ...
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Styracocephalus
''Styracocephalus platyrhynchus'' (Greek for "spiked-head") is an extinct genus of dinocephalian therapsid that existed during the mid-Permian throughout South Africa, but mainly in the Karoo Basin. It is often referred to by its single known species ''Styracocephalus platyrhynchus''. The Dinocephalia clade consisted of the largest land vertebrates and herbivores during the early to mid-Permian. This period is often also referred to as the Guadalupian epoch, approximately 270 to 260 million years ago. Although multiple ''Styracocephalus'' skulls have been recovered, there has yet to be a specimen found of the entire therapsid skeleton. A majority of the skulls collected have also been mature skulls, as the juvenile tapinocephalid skulls are identified by having a small non-fused basioccipital. The presence of enlarged canines, Pachyostosis, pachyostotic cranial boss, and horns are all plesiomorphic traits found in Dinocephalia. ''Styracocephalus''s head Biological ornament, or ...
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Permian Tetrapods
Permian tetrapods were amphibians and reptiles that lived during the Permian Period. During this time, amphibians remained common, including various Temnospondyli and Lepospondyli. Synapsids became the dominant type of animal, represented by the Pelycosaurs during the Early Permian and Therapsids during the Middle and Late Permian, and distinguished by the appearance and possession of mammal-like characteristics (hence the old term "mammal-like reptiles"). These were accompanied by Anapsids or Parareptiles, which included both lizard-like and large herbivorous forms, and primitive diapsids. Classification The following list of families of Permian tetrapods is based mostly on Benton ed. 1993. The classification followBenton 2004 Superclass Tetrapoda * Class Amphibia :::* Order Temnospondyli :::::* Family Edopidae :::::* Family Cochleosauridae :::::* Family Trimerorhachidae :::::* Family Dvinosauridae :::::* Family Saurerpetontidae :::::* Family Brachyopidae :::::* F ...
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Tapinocephalia
The Tapinocephalia are one of the major groups of dinocephalian therapsids and the major herbivorous group. Tapinocephalia has been found to consist of three clades: Styracocephalidae, Titanosuchidae, and the very successful Tapinocephalidae. Notable tapinocephalians include ''Moschops'', ''Tapinocephalus'', and '' Titanosuchus''. Description Unlike anteosaurs and estemmenosuchids, tapinocephalians are primarily an African group. The estemmenosuchids and pareiasaurs may have occupied this paleo-bovine niche in the north. Only one tapinocephalian, ''Ulemosaurus'', is known from Russia. Earlier tapinocephalians were carnivorous or omnivorous. One such group was Titanosuchidae, which consisted of long-tailed predators that hunted herbivorous therapsids Therapsida is a clade comprising a major group of eupelycosaurian synapsids that includes mammals and their ancestors and close relatives. Many of the traits today seen as unique to mammals had their origin within early therap ...
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Estemmenosuchus Mirabilis
''Estemmenosuchus'' (meaning "crowned crocodile" in Greek language, Greek) is an extinct genus of large, early omnivore, omnivorous therapsid. It is believed and interpreted to have lived during the middle part of the Middle Permian around 267 million years ago. The two species, ''E. uralensis'' and ''E. mirabilis'', are characterised by distinctive horn-like structures, which were probably used for display (zoology), intra-specific display. Both species of ''Estemmenosuchus'' are from the Perm, Russia, Perm (or Cis-Urals) region of Russia. Two other estemmenosuchids, ''Anoplosuchus'' and ''Zopherosuchus'', are now considered females of the species ''E. uralensis''. There were many complete and incomplete skeletons found together. Description ''Estemmenosuchus'' could reach a body length of more than . Its skull was long and massive, up to in length, and possessed several sets of large horns, somewhat similar to the antlers of a moose, growing upward and outward from the sides ...
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Gillian King
Gillian may refer to: Places * Gillian Settlement, Arkansas, an unincorporated community People Gillian (variant Jillian) is an English feminine given name, frequently shortened to Gill. It originates as a feminine form of the name Julian, Julio, Julius, and Julien. It is also in use as a surname. Notable people with the name include: First name * Gillian Alexy (born 1986), Australian actress * Gillian Allnutt (born 1949), English poet * Gillian Anderson (born 1968), American actress * Gillian Apps (born 1983), Canadian ice hockey player * Gillian Armstrong (born 1950), Australian film director * Gillian Avery (1926–2016), British children's novelist and literary historian * Gillian Ayres (1930–2018), English painter * Gillian Bailey (born 1955), British academic and actress * Gillian Barge (1940–2003), English actress * Gillian Baverstock (1931–2007), British author * Gillian Baxter (1938–2025), British children's book writer * Gillian Beer (born 1935), Bri ...
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James A
James may refer to: People * James (given name) * James (surname) * James (musician), aka Faruq Mahfuz Anam James, (born 1964), Bollywood musician * James, brother of Jesus * King James (other), various kings named James * Prince James (other) * Saint James (other) Places Canada * James Bay, a large body of water * James, Ontario United Kingdom * James College, York, James College, a college of the University of York United States * James, Georgia, an unincorporated community * James, Iowa, an unincorporated community * James City, North Carolina * James City County, Virginia ** James City (Virginia Company) ** James City Shire * James City, Pennsylvania * St. James City, Florida Film and television * James (2005 film), ''James'' (2005 film), a Bollywood film * James (2008 film), ''James'' (2008 film), an Irish short film * James (2022 film), ''James'' (2022 film), an Indian Kannada-language film * "James", a television Adventure Time (season 5)#ep42, ...
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Basal (phylogenetics)
In phylogenetics, basal is the direction of the ''base'' (or root) of a phylogenetic tree#Rooted tree, rooted phylogenetic tree or cladogram. The term may be more strictly applied only to nodes adjacent to the root, or more loosely applied to nodes regarded as being close to the root. Note that extant taxa that lie on branches connecting directly to the root are not more closely related to the root than any other extant taxa. While there must always be two or more equally "basal" clades sprouting from the root of every cladogram, those clades may differ widely in taxonomic rank, Phylogenetic diversity, species diversity, or both. If ''C'' is a basal clade within ''D'' that has the lowest rank of all basal clades within ''D'', ''C'' may be described as ''the'' basal taxon of that rank within ''D''. The concept of a 'key innovation' implies some degree of correlation between evolutionary innovation and cladogenesis, diversification. However, such a correlation does not make a given ca ...
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Outgroup (cladistics)
In cladistics or phylogenetics, an outgroup is a more distantly related group of organisms that serves as a reference group when determining the evolutionary relationships of the ingroup, the set of organisms under study, and is distinct from sociological outgroups. Character states present in the ingroup but absent in the outgroup are (often) synapomorphies that provide empirical support for the inferred monophyly of the ingroup; character states that are present in the outgroup and some members of the ingroup are symplesiomorphies, and their complementary synapomorphies shared among some members of the ingroup provide hypotheses of relationship within the ingroup clade. The outgroup is used as a point of comparison for the ingroup and specifically allows for the phylogeny to be rooted. Because the polarity (direction) of character change can be determined only on a rooted phylogeny, the choice of outgroup is essential for understanding the evolution of traits along a phylogeny. ...
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Caseidae
Caseidae are an Extinction, extinct Family (biology), family of Basal (phylogenetics), basal synapsids that lived from the Late Carboniferous to Middle Permian between about 300 and 265 million years ago. Fossils of these animals come from the south-central part of the United States (Texas, Oklahoma, and Kansas), from various parts of Europe (European Russia, France, Germany, Sardinia, and Poland), and possibly from South Africa if the genus ''Eunotosaurus'' is indeed a caseid as some authors proposed in 2021. Caseids show great Taxonomy, taxonomic and morphological diversity. The most basal taxa were small Insectivore, insectivorous and Omnivore, omnivorous forms that lived mainly in the Upper Carboniferous and Lower Permian, such as ''Eocasea'', ''Callibrachion'', and ''Martensius''. This type of caseid persists until the Guadalupian, middle Permian with ''Phreatophasma'' and may be ''Eunotosaurus''. During the early Permian, the clade is mainly represented by many species that a ...
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