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Epipubic
Epipubic bones are a pair of bones projecting forward from the pelvic bones of modern marsupials, monotremes and fossil mammals like multituberculates, and even basal eutherians (the ancestors of placentals, who lack them). They first occur in non-mammalian cynodonts such as tritylodontids, suggesting that they are a synapomorphy between them and Mammaliformes. They were first described as early as 1698, but to date, their function(s) remain unresolved. Epipubic bones are often called ''marsupial bones'' because they support the mother's pouch in modern marsupials ("''marsupium''" is Latin for "pouch"). Function Some writers have suggested that the epipubic bones are a part of a kinetic link stretching from the femur on one side, to the ribs on the opposite side. This linkage is formed by a series of muscles: Each epipubic bone is connected to the femur by the pectineus muscle, and to the ribs and vertebrae by the pyramidalis, rectus abdominis, and external and intern ...
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Evolution Of Mammals
The evolution of mammals has passed through many stages since the first appearance of their synapsid ancestors in the Pennsylvanian (geology), Pennsylvanian sub-period of the late Carboniferous period. By the mid-Triassic, there were many synapsid species that looked like mammals. The lineage leading to today's mammals split up in the Jurassic; synapsids from this period include ''Dryolestes'', more closely related to extant placentals and marsupials than to monotremes, as well as ''Ambondro mahabo, Ambondro'', more closely related to monotremes. Later on, the eutherian and metatherian lineages separated; the metatherians are the animals more closely related to the marsupials, while the eutherians are those more closely related to the placentals. Since ''Juramaia'', the earliest known eutherian, lived 160 million years ago in the Jurassic, this divergence must have occurred in the same period. After the Cretaceous–Paleogene extinction event wiped out the non-avian dinosaurs (bi ...
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Placental
Placental mammals (infraclass Placentalia ) are one of the three extant subdivisions of the class Mammalia, the other two being Monotremata and Marsupialia. Placentalia contains the vast majority of extant mammals, which are partly distinguished from monotremes and marsupials in that the fetus (biology), fetus is carried in the uterus of its mother to a relatively late stage of development. The name is something of a misnomer, considering that marsupials also nourish their fetuses via a placenta, though for a relatively briefer period, giving birth to less-developed young, which are then nurtured for a period inside the mother's pouch (marsupial), pouch. Placentalia represents the only living group within Eutheria, which contains all mammals that are more closely related to placentals than they are to marsupials. Anatomical features Placental mammals are anatomically distinguished from other mammals by: * a sufficiently wide opening at the bottom of the pelvis to allow the birt ...
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Mammaliformes
Mammaliaformes ("mammalian forms") is a clade of synapsid tetrapods that includes the crown group mammals and their closest extinct relatives; the group radiated from earlier probainognathian cynodonts during the Late Triassic. It is defined as the clade originating from the most recent common ancestor of Morganucodonta and the crown group mammals; the latter is the clade originating with the most recent common ancestor of extant Monotremata, Marsupialia and Placentalia. Besides Morganucodonta and the crown group mammals, Mammaliaformes also includes Docodonta and ''Hadrocodium''. Mammaliaformes is a term of phylogenetic nomenclature. In contrast, the assignment of organisms to class Mammalia has traditionally been founded on traits and, on this basis, Mammalia is slightly more inclusive than Mammaliaformes. In particular, trait-based taxonomy generally includes ''Adelobasileus'' and '' Sinoconodon'' in Mammalia, though they fall outside the Mammaliaformes definition. These ge ...
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Eutheria
Eutheria (from Greek , 'good, right' and , 'beast'; ), also called Pan-Placentalia, is the clade consisting of Placentalia, placental mammals and all therian mammals that are more closely related to placentals than to marsupials. Eutherians are distinguished from non-eutherians by various phenotypic traits of the feet, ankles, jaws and teeth. All extant eutherians lack epipubic bones, which are present in all other living mammals (marsupials and monotremes). This allows for expansion of the abdomen during pregnancy, though epipubic bones are present in many primitive eutherians. Eutheria was named in 1872 by Theodore Gill; in 1880, Thomas Henry Huxley defined it to encompass a more broadly defined group than Placentalia. The earliest unambiguous eutherians are known from the Early Cretaceous Yixian Formation of China, dating around 120 million years ago. Two tribosphenic mammals, ''Durlstodon'' and ''Durlstotherium'' from the Berriasian age (~145–140 million years ago) of t ...
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Multituberculate
Multituberculata (commonly known as multituberculates, named for the multiple tubercles of their teeth) is an extinct order of rodent-like mammals with a fossil record spanning over 130 million years. They first appeared in the Middle Jurassic, and reached a peak diversity during the Late Cretaceous and Paleocene. They eventually declined from the mid-Paleocene onwards, disappearing from the known fossil record in the late Eocene. They are the most diverse order of Mesozoic mammals with more than 200 species known, ranging from mouse-sized to beaver-sized. These species occupied a diversity of ecological niches, ranging from burrow-dwelling to squirrel-like arborealism to jerboa-like hoppers. Multituberculates are usually placed as crown mammals outside either of the two main groups of living mammals, Theria — placentals and marsupials — and Monotremata,Agustí-Antón 2002, pp 3-4 but usually as closer to Theria than to monotremes. They are considered to be closely re ...
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Baculum
The baculum (: bacula), also known as the penis bone, penile bone, ''os penis'', ''os genitale'', or ''os priapi'', is a bone in the penis of many placental mammals. It is not present in humans, but is present in the penises of some primates, such as the gorilla and the chimpanzee. The baculum arises from primordial cells in soft tissues of the penis, and its formation is largely influenced by androgens. The bone lies above the male urethra, urethra, and it aids sexual reproduction by maintaining stiffness during sexual penetration. The Homology (biology), homologue to the baculum in female mammals is the baubellum (''os clitoridis''), a bone in the clitoris. Etymology The word ''baculum'' means "stick" or "staff" in Latin and originates from , ''baklon'' "stick". Function The baculum is used for copulation and varies in size and shape by species. Its evolution may be influenced by sexual selection, and its characteristics are sometimes used to differentiate between simila ...
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Kayentatherium
''Kayentatherium'' is an extinct genus of tritylodontid cynodonts that lived during the Early Jurassic. It is one of two tritylodonts from the Kayenta Formation of northern Arizona, United States. ''Kayentatherium'' means "Kayenta Beast", and is named for the geological formation in which it was found, the Kayenta Formation. ''Kayentatherium'' is known from several specimens.Kermack, D. 1982. A new tritylodont from the Kayenta Formation of Arizona. Zoological Journal of the Linnean Society, 76. 1-17.Sues, Hans-Dieter & F. A. Jenkins. 2006. The Postcranial Skeleton of Kayentatherium Wellesi from the Lower Jurassic Kayenta Formation of Arizona and the Phylogenetic Significance of Postcranial Features in Tritylodontid Cynodonts in: Carrano, Matthew T., Gaudin, T. J., Blob, R. W. and Wible, J. R., Amniote Paleobiology: Perspectives on the Evolution of Mammals, Birds, and Reptiles: The University of Chicago Press. pp. 114-152. The specific name honors paleontologist Samuel Welles, ...
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Placenta
The placenta (: placentas or placentae) is a temporary embryonic and later fetal organ that begins developing from the blastocyst shortly after implantation. It plays critical roles in facilitating nutrient, gas, and waste exchange between the physically separate maternal and fetal circulations, and is an important endocrine organ, producing hormones that regulate both maternal and fetal physiology during pregnancy. The placenta connects to the fetus via the umbilical cord, and on the opposite aspect to the maternal uterus in a species-dependent manner. In humans, a thin layer of maternal decidual ( endometrial) tissue comes away with the placenta when it is expelled from the uterus following birth (sometimes incorrectly referred to as the 'maternal part' of the placenta). Placentas are a defining characteristic of placental mammals, but are also found in marsupials and some non-mammals with varying levels of development. Mammalian placentas probably first evolved abou ...
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Flexion
Motion, the process of movement, is described using specific anatomical terminology, anatomical terms. Motion includes movement of Organ (anatomy), organs, joints, Limb (anatomy), limbs, and specific sections of the body. The terminology used describes this motion according to its direction relative to the anatomical position of the body parts involved. Anatomy, Anatomists and others use a unified set of terms to describe most of the movements, although other, more specialized terms are necessary for describing unique movements such as those of the hands, feet, and eyes. In general, motion is classified according to the anatomical plane it occurs in. ''Flexion'' and ''extension'' are examples of ''angular'' motions, in which two axes of a joint are brought closer together or moved further apart. ''Rotational'' motion may occur at other joints, for example the shoulder, and are described as ''internal'' or ''external''. Other terms, such as ''elevation'' and ''depression'', descri ...
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Abdominal Internal Oblique Muscle
The abdominal internal oblique muscle, also internal oblique muscle or interior oblique, is an abdominal muscle in the abdominal wall that lies below the external oblique muscle and just above the transverse abdominal muscle. Structure Its fibers run perpendicular to the external oblique muscle, beginning in the thoracolumbar fascia of the lower back, the anterior 2/3 of the iliac crest (upper part of hip bone) and the lateral half of the inguinal ligament. The muscle fibers run from these points superomedially (up and towards midline) to the muscle's insertions on the inferior borders of the 10th through 12th ribs and the linea alba. In males, the cremaster muscle is also attached to the internal oblique. Nerve supply The internal oblique is supplied by the lower intercostal nerves, as well as the iliohypogastric nerve and the ilioinguinal nerve. Function The internal oblique performs two major functions. Firstly as an accessory muscle of respiration, it acts as an ...
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Respiration (physiology)
In physiology, respiration is the transport of oxygen from the outside environment to the Cell (biology), cells within Tissue (biology), tissues, and the removal of carbon dioxide in the opposite direction to the environment by a respiratory system. The physiological definition of respiration differs from the Cellular respiration, biochemical definition, which refers to a metabolic process by which an organism obtains energy (in the form of ATP and NADPH) by oxidizing nutrients and releasing waste products. Although physiologic respiration is necessary to sustain cellular respiration and thus life in animals, the processes are distinct: cellular respiration takes place in individual cells of the organism, while physiologic respiration concerns the Diffusion#Diffusion vs. bulk flow diffusion, diffusion and transport of metabolites between the organism and the external environment. Exchange of gases in the lung occurs by ventilation and perfusion. Ventilation refers to the in-and-ou ...
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Hypaxial
In adult vertebrates, trunk muscles can be broadly divided into hypaxial muscles, which lie ventral to the horizontal septum of the vertebrae and epaxial muscles, which lie dorsal to the septum. Hypaxial muscles include some vertebral muscles, the diaphragm, the abdominal muscles, and all limb muscles. The serratus posterior inferior and serratus posterior superior are innervated by the ventral primary ramus and are hypaxial muscles. Epaxial muscles include other (dorsal) muscles associated with the vertebrae, ribs, and base of the skull. In humans, the erector spinae, the transversospinales (including the multifidus, semispinalis and rotatores), the splenius and suboccipital muscles are the only epaxial muscles. Hypaxial and epaxial muscles develop directly from somitic cells. Differentiation of hypaxial and epaxial muscles is postulated to have evolved as a new trait in vertebrate animals. Location The hypaxial muscles are located on the ventral side of the body, ...
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