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Epiaceratherium
''Epiaceratherium'' is an extinct genus of rhinocerotid from the Eocene and Oligocene of Europe, Asia, and North America. Taxonomy The genus was named by paleontologist Othenio Abel in 1910, with the type species being ''Epiaceratherium bolcense.'' This species is exclusively known from remains found at Monteviale in northern Italy, dating to the earliest Oligocene (~34 million years ago). The species ''Epiaceratherium magnum'' named by Uhlig, 1999, is known from remains found in Germany, France, Czechia and Switzerland, dating to the Early Oligocene to early Late Oligocene. Remains similar to this species have also been reported from Pakistan, dating to the Early Oligocene. In 2013 the species ''Epiaceratherium naduongense'' was described from Na Duong Basin in northern Vietnam, dating to the mid-late Eocene (~39–35 million years ago). In 2021, the species ''Molassitherium delemontense'' originally described in 2013 from late Early–early Late Oligocene deposits in Germany, Swit ...
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Rhinocerotidae
A rhinoceros ( ; ; ; : rhinoceros or rhinoceroses), commonly abbreviated to rhino, is a member of any of the five extant taxon, extant species (or numerous extinct species) of odd-toed ungulates (perissodactyls) in the family (biology), family Rhinocerotidae; it can also refer to a member of any of the extinct species of the superfamily Rhinocerotoidea. Two of the extant species are native to Africa, and three to South Asia, South and Southeast Asia. Rhinoceroses are some of the largest remaining megafauna: all weigh over half a tonne in adulthood. They have a herbivore, herbivorous diet, small brains for mammals of their size, one or two horns, and a thick , protective skin formed from layers of collagen positioned in a crystal structure, lattice structure. They generally eat leafy material, although their ability to ferment food in their colon (anatomy), hindgut allows them to subsist on more fibrous plant matter when necessary. Unlike other perissodactyls, the two African ...
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Rhinocerotinae
A rhinoceros ( ; ; ; : rhinoceros or rhinoceroses), commonly abbreviated to rhino, is a member of any of the five extant species (or numerous extinct species) of odd-toed ungulates (perissodactyls) in the family Rhinocerotidae; it can also refer to a member of any of the extinct species of the superfamily Rhinocerotoidea. Two of the extant species are native to Africa, and three to South and Southeast Asia. Rhinoceroses are some of the largest remaining megafauna: all weigh over half a tonne in adulthood. They have a herbivorous diet, small brains for mammals of their size, one or two horns, and a thick , protective skin formed from layers of collagen positioned in a lattice structure. They generally eat leafy material, although their ability to ferment food in their hindgut allows them to subsist on more fibrous plant matter when necessary. Unlike other perissodactyls, the two African species of rhinoceros lack teeth at the front of their mouths; they rely instead on their l ...
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Elasmotheriinae
A rhinoceros ( ; ; ; : rhinoceros or rhinoceroses), commonly abbreviated to rhino, is a member of any of the five extant taxon, extant species (or numerous extinct species) of odd-toed ungulates (perissodactyls) in the family (biology), family Rhinocerotidae; it can also refer to a member of any of the extinct species of the superfamily Rhinocerotoidea. Two of the extant species are native to Africa, and three to South Asia, South and Southeast Asia. Rhinoceroses are some of the largest remaining megafauna: all weigh over half a tonne in adulthood. They have a herbivore, herbivorous diet, small brains for mammals of their size, one or two horns, and a thick , protective skin formed from layers of collagen positioned in a crystal structure, lattice structure. They generally eat leafy material, although their ability to ferment food in their colon (anatomy), hindgut allows them to subsist on more fibrous plant matter when necessary. Unlike other perissodactyls, the two African ...
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Trigonias
''Trigonias'' (Greek: "triangular" (trigonos), "ias" enotes possession is an extinct genus of rhinocerotid from the late Eocene (Chadronian) some 35 million years ago of North America. ''Trigonias'' was about long and, despite lacking horns, looked a lot like modern rhinos. Its front legs had five toes (as contrasted with three in modern rhinos), the fifth of which was vestigial Vestigiality is the retention, during the process of evolution, of genetically determined structures or attributes that have lost some or all of the ancestral function in a given species. Assessment of the vestigiality must generally rely on co .... A specimen of ''T. osborni'' was estimated to have a weight of about .Paleobiology Database
Cladogram after Lu, Deng and Pandolfi, 2023:


Notes



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Oligocene Rhinoceroses
The Oligocene ( ) is a geologic epoch of the Paleogene Period that extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but the exact dates of the start and end of the epoch are slightly uncertain. The name Oligocene was coined in 1854 by the German paleontologist Heinrich Ernst Beyrich from his studies of marine beds in Belgium and Germany. The name comes from Ancient Greek (''olígos'') 'few' and (''kainós'') 'new', and refers to the sparsity of extant forms of molluscs. The Oligocene is preceded by the Eocene Epoch and is followed by the Miocene Epoch. The Oligocene is the third and final epoch of the Paleogene Period. The Oligocene is often considered an important time of transition, a link between the archaic world of the tropical Eocene and the more modern ecosystems of the Miocene. Major changes during the Oligocene included a global expansion of gra ...
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Molar (tooth)
The molars or molar teeth are large, flat teeth at the back of the mouth. They are more developed in mammals. They are used primarily to grind food during chewing. The name ''molar'' derives from Latin, ''molaris dens'', meaning "millstone tooth", from ''mola'', millstone and ''dens'', tooth. Molars show a great deal of diversity in size and shape across the mammal groups. The third molar of humans is sometimes vestigial. Human anatomy In humans, the molar teeth have either four or five cusps. Adult humans have 12 molars, in four groups of three at the back of the mouth. The third, rearmost molar in each group is called a wisdom tooth. It is the last tooth to appear, breaking through the front of the gum at about the age of 20, although this varies among individuals and populations, and in many cases the tooth is missing. The human mouth contains upper (maxillary) and lower (mandibular) molars. They are: maxillary first molar, maxillary second molar, maxillary third mol ...
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Premolar
The premolars, also called premolar Tooth (human), teeth, or bicuspids, are transitional teeth located between the Canine tooth, canine and Molar (tooth), molar teeth. In humans, there are two premolars per dental terminology#Quadrant, quadrant in the permanent teeth, permanent set of teeth, making eight premolars total in the mouth. They have at least two Cusp (dentistry), cusps. Premolars can be considered transitional teeth during chewing, or mastication. They have properties of both the canines, that lie anterior and molars that lie Posterior (anatomy), posterior, and so food can be transferred from the canines to the premolars and finally to the molars for grinding, instead of directly from the canines to the molars. Human anatomy The premolars in humans are the maxillary first premolar, maxillary second premolar, mandibular first premolar, and the mandibular second premolar. Premolar teeth by definition are permanent teeth Anatomical terms of location#Proximal and distal, ...
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Canine Tooth
In mammalian oral anatomy, the canine teeth, also called cuspids, dogteeth, eye teeth, vampire teeth, or fangs, are the relatively long, pointed teeth. In the context of the upper jaw, they are also known as '' fangs''. They can appear more flattened, however, causing them to resemble incisors and leading them to be called ''incisiform''. They developed and are used primarily for firmly holding food in order to tear it apart, and occasionally as weapons. They are often the largest teeth in a mammal's mouth. Individuals of most species that develop them normally have four, two in the upper jaw and two in the lower, separated within each jaw by incisors; humans and dogs are examples. In most species, canines are the anterior-most teeth in the maxillary bone. The four canines in humans are the two upper maxillary canines and the two lower mandibular canines. They are specially prominent in dogs (Canidae), hence the name. Details There are generally four canine teeth: two ...
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Incisor
Incisors (from Latin ''incidere'', "to cut") are the front teeth present in most mammals. They are located in the premaxilla above and on the mandible below. Humans have a total of eight (two on each side, top and bottom). Opossums have 18, whereas armadillos, anteaters and other animals in the order Edentata have none. Structure Adult humans normally have eight incisors, two of each type. The types of incisors are: * maxillary central incisor (upper jaw, closest to the center of the lips) * maxillary lateral incisor (upper jaw, beside the maxillary central incisor) * mandibular central incisor (lower jaw, closest to the center of the lips) * mandibular lateral incisor (lower jaw, beside the mandibular central incisor) Children with a full set of deciduous teeth (primary teeth) also have eight incisors, named the same way as in permanent teeth. Young children may have from zero to eight incisors depending on the stage of their tooth eruption and tooth development. Typic ...
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Menoceras
''Menoceras'' ("Crescent Horns") is a genus of extinct, small rhinocerotids endemic to most of southern North America and ranged as far south as Panama during the early Miocene epoch. It lived from around 23.1-12.5 Ma, existing for approximately . Description ''Menoceras'' was much smaller than any living rhinoceros, with the genus being compared in size to a sheep or a pig,D.R. Prother"Rhinocerotidae"C.M. Janis, K.M. Scott, L. Jacobs (Eds.), Evolution of Tertiary Mammals of North America, Cambridge University Press, Cambridge (1998), pp. 595-605 an with estimated bodymass of . Male ''Menoceras'' sported two horns side by side at the tip of the nose, whereas the females were hornless or had greatly reduced horns, and have differently shaped nasal bones. ''Menoceras'' along with '' Diceratherium'' is unique in having paired horns among rhinocerotids. Both sexes of ''Menoceros'' grew to a length of long. Paleobiology ''Menoceras'' roamed across a tropical, savanna-like grassland ...
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Protaceratherium
''Protaceratherium'' is an extinct genus of rhinocerotid from the Oligocene and Miocene of Eurasia. It was a primitive, lightly built rhinoceros that was adapted to running. Palaeoecology Analysis of dental Δ13C, δ13C values combined with dental mesowear and microwear show that ''P. minutum'' fed primarily on C3 plants, C3 plants and that it consumed less abrasive plants than the contemporary ''Mesaceratherium, Mesaceratherium paulhiacense''. References

Oligocene rhinoceroses Oligocene mammals of Europe Oligocene mammals of Asia Miocene rhinoceroses Miocene mammals of Europe Miocene mammals of Asia {{paleo-oddtoedungulate-stub ...
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