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Entodacrya
The term Hystricomorpha (from Greek ὕστριξ, ''hystrix'' 'porcupine' and Greek μορφή, ''morphē'' 'form') has had many definitions throughout its history. In the broadest sense, it refers to any rodent (except dipodoids) with a hystricomorphous zygomasseteric system. This includes the Hystricognathi, Ctenodactylidae, Anomaluridae, and Pedetidae. Molecular and morphological results suggest the inclusion of the Anomaluridae and Pedetidae in Hystricomorpha may be suspect. Based on , these two families are discussed here as representing a distinct suborder Anomaluromorpha. Classification The modern definition of Hystricomorpha, also known as Entodacrya or Ctenohystrica, is a taxonomic hypothesis uniting the gundis with the hystricognath rodents. Considerable morphological and strong molecular support exists for this relationship. If true, this hypothesis renders the traditional view of Sciurognathi invalid, as it becomes a paraphyletic group. The hystricomorph roden ...
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Eocene
The Eocene ( ) Epoch is a geological epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', " dawn") and (''kainós'', "new") and refers to the "dawn" of modern ('new') fauna that appeared during the epoch. The Eocene spans the time from the end of the Paleocene Epoch to the beginning of the Oligocene Epoch. The start of the Eocene is marked by a brief period in which the concentration of the carbon isotope 13C in the atmosphere was exceptionally low in comparison with the more common isotope 12C. The end is set at a major extinction event called the ''Grande Coupure'' (the "Great Break" in continuity) or the Eocene–Oligocene extinction event, which may be related to the impact of one or more large bolides in Siberia and in what is now Chesapeake Bay. As with other geologic periods, the strata that define the start and e ...
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Caviomorpha
Caviomorpha is the rodent infraorder or parvorder that unites all New World hystricognaths. It is supported by both fossil and molecular evidence. The Caviomorpha was for a time considered to be a separate order outside the Rodentia, but is now accepted as a genuine part of the rodents. Caviomorphs include the extinct Heptaxodontidae (giant hutias) and extant families of chinchilla rats, hutias, guinea pigs and the capybara, chinchillas and viscachas, tuco-tucos, agoutis, pacas, pacaranas, spiny rats, New World porcupines, coypu and octodonts (Vassallo and Antenucci, 2015). Origin The first known rodent fossils in South America are represented by the three taxa ''Cachiyacuy contamanensis'', ''C. kummeli'', and ''Canaanimys maquiensis'', as well as teeth from ''Eobranisamys'' sp. (Dasyproctidae) and ''Eospina'' sp., the latter two found also in the Santa Rosa fauna from the late Eocene or early Oligocene. By the late Oligocene, all superfamilies and most families of cavio ...
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Taxonomy (biology)
In biology, taxonomy () is the scientific study of naming, defining ( circumscribing) and classifying groups of biological organisms based on shared characteristics. Organisms are grouped into taxa (singular: taxon) and these groups are given a taxonomic rank; groups of a given rank can be aggregated to form a more inclusive group of higher rank, thus creating a taxonomic hierarchy. The principal ranks in modern use are domain, kingdom, phylum (''division'' is sometimes used in botany in place of ''phylum''), class, order, family, genus, and species. The Swedish botanist Carl Linnaeus is regarded as the founder of the current system of taxonomy, as he developed a ranked system known as Linnaean taxonomy for categorizing organisms and binomial nomenclature for naming organisms. With advances in the theory, data and analytical technology of biological systematics, the Linnaean system has transformed into a system of modern biological classification intended to reflect the ...
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Great American Interchange
The Great American Biotic Interchange (commonly abbreviated as GABI), also known as the Great American Interchange and the Great American Faunal Interchange, was an important late Cenozoic paleozoogeographic biotic interchange event in which land and freshwater fauna migrated from North America via Central America to South America and vice versa, as the volcanic Isthmus of Panama rose up from the sea floor and bridged the formerly separated continents. Although earlier dispersals had occurred, probably over water, the migration accelerated dramatically about 2.7 million years ( Ma) ago during the Piacenzian age. It resulted in the joining of the Neotropic (roughly South American) and Nearctic (roughly North American) biogeographic realms definitively to form the Americas. The interchange is visible from observation of both biostratigraphy and nature ( neontology). Its most dramatic effect is on the zoogeography of mammals, but it also gave an opportunity for reptiles, ...
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Primates
Primates are a diverse order of mammals. They are divided into the strepsirrhines, which include the lemurs, galagos, and lorisids, and the haplorhines, which include the tarsiers and the simians (monkeys and apes, the latter including humans). Primates arose 85–55 million years ago first from small terrestrial mammals, which adapted to living in the trees of tropical forests: many primate characteristics represent adaptations to life in this challenging environment, including large brains, visual acuity, color vision, a shoulder girdle allowing a large degree of movement in the shoulder joint, and dextrous hands. Primates range in size from Madame Berthe's mouse lemur, which weighs , to the eastern gorilla, weighing over . There are 376–524 species of living primates, depending on which classification is used. New primate species continue to be discovered: over 25 species were described in the 2000s, 36 in the 2010s, and three in the 2020s. Primates hav ...
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Africa
Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area and 20% of its land area.Sayre, April Pulley (1999), ''Africa'', Twenty-First Century Books. . With billion people as of , it accounts for about of the world's human population. Africa's population is the youngest amongst all the continents; the median age in 2012 was 19.7, when the worldwide median age was 30.4. Despite a wide range of natural resources, Africa is the least wealthy continent per capita and second-least wealthy by total wealth, behind Oceania. Scholars have attributed this to different factors including geography, climate, tribalism, Scramble for Africa, colonialism, the Cold War, neocolonialism, lack of democracy, and corruption. Despite this low concentration of wealth, recent economic expansion and the large and young ...
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Atlantic Ocean
The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about . It covers approximately 20% of Earth#Surface, Earth's surface and about 29% of its water surface area. It is known to separate the "Old World" of Africa, Europe and Asia from the "New World" of the Americas in the European perception of Earth, the World. The Atlantic Ocean occupies an elongated, S-shaped basin extending longitudinally between Europe and Africa to the east, and North America, North and South America to the west. As one component of the interconnected World Ocean, it is connected in the north to the Arctic Ocean, to the Pacific Ocean in the southwest, the Indian Ocean in the southeast, and the Southern Ocean in the south (other definitions describe the Atlantic as extending southward to Antarctica). The Atlantic Ocean is divided in two parts, by the Equatorial Counter Current, with the North(ern) Atlantic Ocean and the South(ern) Atlantic Ocean split at about 8th paralle ...
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Oceanic Dispersal
Oceanic dispersal is a type of biological dispersal that occurs when terrestrial organisms transfer from one land mass to another by way of a sea crossing. Island hopping is the crossing of an ocean by a series of shorter journeys between islands, as opposed to a single journey directly to the destination. Often this occurs via large rafts of floating vegetation such as are sometimes seen floating down major rivers in the tropics and washing out to sea, occasionally with animals trapped on them. Dispersal via such a raft is sometimes referred to as a rafting event. Colonization of land masses by plants can also occur via long-distance oceanic dispersal of floating seeds. History Rafting has played an important role in the colonization of isolated land masses by mammals. Prominent examples include Madagascar, which has been isolated for ~120 million years ( Ma), and South America, which was isolated for much of the Cenozoic. Both land masses, for example, appear to have receiv ...
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Meridiungulata
South American native ungulates, commonly abbreviated as SANUs, are extinct ungulate-like mammals of controversial affinities that were indigenous to South America prior to the Great American Biotic Interchange. They comprise five major groups conventionally ranked as orders— Astrapotheria, Litopterna, Notoungulata, Pyrotheria, and Xenungulata—as well as some other taxa, such as Didolodontidae and Kollpaniidae. It has been proposed that some or all of the members of this group form a clade, named Meridiungulata, though the relationships of South American ungulates remain largely unresolved. The two largest groups of South American ungulates, the notoungulates and the litopterns, were the only groups to persist beyond the mid Miocene. Only a few of the largest species of notoungulates and litopterns survived until the end-Pleistocene extinctions. Though most SANUs lived in South America, astrapotheres and litopterns are known from Eocene aged deposits in the Antarctic Pe ...
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Xenarthra
Xenarthra (; from Ancient Greek ξένος, xénos, "foreign, alien" + ἄρθρον, árthron, "joint") is a major clade of placental mammals native to the Americas. There are 31 living species: the anteaters, tree sloths, and armadillos. Extinct xenarthrans include the glyptodonts, pampatheres and ground sloths. Xenarthrans originated in South America during the late Paleocene about 60 million years ago. They evolved and diversified extensively in South America during the continent's long period of isolation in the early to mid Cenozoic Era. They spread to the Antilles by the early Miocene and, starting about 3 Mya, spread to Central and North America as part of the Great American Interchange. Nearly all of the formerly abundant megafaunal xenarthrans became extinct at the end of the Pleistocene. Xenarthrans share several characteristics not present in other placental mammals, which suggest their ancestors were subterranean diggers for insects. The name Xenarthra deriv ...
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Metatheria
Metatheria is a mammalian clade that includes all mammals more closely related to marsupials than to placentals. First proposed by Thomas Henry Huxley in 1880, it is a more inclusive group than the marsupials; it contains all marsupials as well as many extinct non-marsupial relatives. There are three extant subclasses of mammals, one being metatherians: # monotremes: egg laying mammals like the platypus and the echidna, #metatheria: marsupials, which includes three American orders ( Didelphimorphia, Paucituberculata and Microbiotheria) and four Australasian orders ( Notoryctemorphia, Dasyuromorphia, Peramelemorphia and Diprotodontia), and the #eutherians: placental mammals, consisting of four superorders divided into 21 orders. Metatherians belong to a subgroup of the northern tribosphenic mammal clade or Boreosphenida. They differ from all other mammals in certain morphologies like their dental formula, which includes about five upper and four lower incisors, a canine ...
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South America
South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southern Subregion#Americas, subregion of a single continent called Americas, America. South America is bordered on the west by the Pacific Ocean and on the north and east by the Atlantic Ocean; North America and the Caribbean Sea lie to the northwest. The continent generally includes twelve sovereign states: Argentina, Bolivia, Brazil, Chile, Colombia, Ecuador, Guyana, Paraguay, Peru, Suriname, Uruguay, and Venezuela; two dependent territory, dependent territories: the Falkland Islands and South Georgia and the South Sandwich Islands; and one administrative division, internal territory: French Guiana. In addition, the ABC islands (Leeward Antilles), ABC islands of the Kingdom of the Netherlands, Ascension Island (dependency of Saint Helena, Asce ...
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