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Elmisauridae
Caenagnathidae is a family of derived caenagnathoid dinosaurs from the Cretaceous of North America and Asia. They are a member of the Oviraptorosauria, and relatives of the Oviraptoridae. Like other oviraptorosaurs, caenagnathids had specialized beaks, long necks, and short tails, and would have been covered in feathers. The relationships of caenagnathids were long a puzzle. The family was originally named by Raymond Martin Sternberg in 1940 as a family of flightless birds. The discovery of skeletons of the related oviraptorids revealed that they were in fact non-avian theropods, and the discovery of more complete caenagnathid remains revealed that ''Chirostenotes pergracilis'', originally named on the basis of a pair of hands, and ''Citipes elegans'', originally thought to be an ornithomimid, named from a foot, were caenagnathids as well. Discovery The name ''Caenagnathus'' (and hence Caenagnathidae) means "recent jaws"—when first discovered, it was thought that caenagnathid ...
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Halszka Osmólska
Halszka Osmólska (September 15, 1930 – March 31, 2008) was a Polish paleontologist who had specialized in Mongolian dinosaurs. Early life, family and education Osmólska was born in 1930 in Poznań. In 1949, she began to study biology at Faculty of Biology and Earth Sciences of the University of Poznań. Thereafter, she studied at the University of Warsaw, graduating in 1955. Career Osmólska worked at the Institute of Paleobiology of the Polish Academy of Sciences (PAN). Between 1983 and 1988, she served as the institute's director. She was a member of the Polish–Mongolian expeditions to the Gobi Desert (1963–1965, 1967–1971), and she described many finds from these rocks, often with Teresa Maryańska. Among the dinosaurs she described: * '' Deinocheirus'' (1967), with Ewa Roniewicz * '' Gallimimus'' (1972), with Roniewicz and Rinchen Barsbold * '' Pachycephalosauria'' (1974), with Maryańska * ''Bagaceratops'' (1975), with Maryańska * '' Barsboldia'' (1981) ...
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Oviraptoridae
Oviraptoridae is a group of bird-like, herbivorous and omnivorous maniraptoran dinosaurs. Oviraptorids are characterized by their toothless, parrot-like beaks and, in some cases, elaborate crests. They were generally small, measuring between one and two metres long in most cases, though some possible oviraptorids were enormous. Oviraptorids are currently known only from the Late Cretaceous of Asia, with the most well-known species and complete specimens found only in the Gobi Desert of Mongolia and northwestern China. Description The most characteristic feature of this group is the skull structure. Oviraptorids had short snouts and very deep mandibles. Some taxa (such as ''Citipati'', '' Corythoraptor'', ''Rinchenia'') had a midline crest on top of the skull, resembling that of a cassowary. Other distinguishing characteristics include a bony spike intruding on the mandibular fenestra, nostrils placed very high and far back on the snout, an extremely thin bony bar beneath the e ...
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Early Cretaceous
The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name) is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 143.1 Megaannum#SI prefix multipliers, Ma to 100.5 Ma. Geology Proposals for the exact age of the Barremian–Aptian boundary ranged from 126 to 117 Ma until recently (as of 2019), but based on drillholes in Svalbard the defining Anoxic event#Cretaceous, early Aptian Oceanic Anoxic Event 1a (OAE1a) was dated to 123.1±0.3 Ma, limiting the possible range for the boundary to c. 122–121 Ma. There is a possible link between this anoxic event and a series of Early Cretaceous large igneous provinces (LIP). The Ontong Java Plateau, Ontong Java-Manihiki Plateau, Manihiki-Hikurangi Plateau, Hikurangi large igneous province, emplaced in the South Pacific at c. 120 Ma, is by far the largest LIP in Earth's history. The Onto ...
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Caenagnathasia
''Caenagnathasia'' ('recent jaw from Asia') is a small caenagnathid oviraptorosaurian theropod from the Late Cretaceous of Uzbekistan. Discovery The type species ''Caenagnathasia martinsoni'' was named and described in 1994 by Philip J. Currie, Stephen Godfrey and Lev Nesov. The generic name is a combination of a reference to the species' placement in the Caenagnathidae, with Asia, the continent of its provenance. The specific name honours Gerbert Genrikhovich Martinson. The genus is based on holotype N 401/12457, a pair of fused dentaries of the lower jaws. The specimen was found near Dzharakuduk in layers of the Bissekty Formation, dating to the Turonian-Coniacian, around 90 mya, making it the oldest known caenagnathoid. A second specimen was referred to the species, N 402/12457, a right dentary of a slightly smaller individual. Both individuals were adult. In 2015, new material of ''Caenagnathasia'' was described. From the same site as the holotype, the materi ...
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Paleognathae
Palaeognathae (; ) is an infraclass of birds, called paleognaths or palaeognaths, within the class Aves of the clade Archosauria. It is one of the two extant infraclasses of birds, the other being Neognathae, both of which form Neornithes. Palaeognathae contains five extant orders consisting of four flightless lineages (plus two that are extinct), termed ratites, and one flying lineage, the Neotropic tinamous. There are 47 species of tinamous, five of kiwis (''Apteryx''), three of cassowaries (''Casuarius''), one of emus (''Dromaius'') (another became extinct in historic times), two of rheas (''Rhea'') and two of ostriches ('' Struthio'').Clements, J. C. ''et al''. (2010) Recent research has indicated that paleognaths are monophyletic but the traditional taxonomic split between flightless and flighted forms is incorrect; tinamous are within the ratite radiation, meaning flightlessness arose independently multiple times via parallel evolution. There are three extinct groups t ...
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Oviraptorosauria
Oviraptorosaurs ("egg thief lizards") are a group of feathered maniraptoran dinosaurs from the Cretaceous Period (geology), Period of what are now Asia and North America. They are distinct for their characteristically short, beaked, parrot-like skulls, with or without bony crests atop the head. They ranged in size from ''Caudipteryx'', which was the size of a turkey, to the 8-meter-long, 1.4-ton ''Gigantoraptor''. The group (along with all maniraptoran dinosaurs) is close to the Origin of birds, ancestry of birds. Some researchers such as Teresa Maryanska, Maryanska ''et al'' (2002) and Halszka Osmólska, Osmólska ''et al.'' (2004) have proposed that they may represent primitive flightless birds.Osmólska, Halszka, Currie, Philip J., Brasbold, Rinchen (2004) "The Dinosauria" Weishampel, Dodson, Osmólska. "Chapter 8 Oviraptorosauria" University of California Press. The most complete oviraptorosaur specimens have been found in Asia. The North American oviraptorosaur record is spa ...
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Asia
Asia ( , ) is the largest continent in the world by both land area and population. It covers an area of more than 44 million square kilometres, about 30% of Earth's total land area and 8% of Earth's total surface area. The continent, which has long been home to the majority of the human population, was the site of many of the first civilisations. Its 4.7 billion people constitute roughly 60% of the world's population. Asia shares the landmass of Eurasia with Europe, and of Afro-Eurasia with both Europe and Africa. In general terms, it is bounded on the east by the Pacific Ocean, on the south by the Indian Ocean, and on the north by the Arctic Ocean. The border of Asia with Europe is a social constructionism, historical and cultural construct, as there is no clear physical and geographical separation between them. A commonly accepted division places Asia to the east of the Suez Canal separating it from Africa; and to the east of the Turkish straits, the Ural Mountains an ...
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North America
North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Caribbean Sea, and to the south and west by the Pacific Ocean. The region includes Middle America (Americas), Middle America (comprising the Caribbean, Central America, and Mexico) and Northern America. North America covers an area of about , representing approximately 16.5% of Earth's land area and 4.8% of its total surface area. It is the third-largest continent by size after Asia and Africa, and the list of continents and continental subregions by population, fourth-largest continent by population after Asia, Africa, and Europe. , North America's population was estimated as over 592 million people in list of sovereign states and dependent territories in North America, 23 independent states, or about 7.5% of the world's popula ...
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Cretaceous
The Cretaceous ( ) is a geological period that lasted from about 143.1 to 66 mya (unit), million years ago (Mya). It is the third and final period of the Mesozoic Era (geology), Era, as well as the longest. At around 77.1 million years, it is the ninth and longest geological period of the entire Phanerozoic. The name is derived from the Latin , 'chalk', which is abundant in the latter half of the period. It is usually abbreviated K, for its German translation . The Cretaceous was a period with a relatively warm climate, resulting in high Sea level#Local and eustatic, eustatic sea levels that created numerous shallow Inland sea (geology), inland seas. These oceans and seas were populated with now-extinct marine reptiles, ammonites, and rudists, while dinosaurs continued to dominate on land. The world was largely ice-free, although there is some evidence of brief periods of glaciation during the cooler first half, and forests extended to the poles. Many of the dominant taxonomic gr ...
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Dinosaur
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutionary history, evolution of dinosaurs is a subject of active research. They became the dominant terrestrial vertebrates after the Triassic–Jurassic extinction event 201.3 mya and their dominance continued throughout the Jurassic and Cretaceous periods. The fossil record shows that birds are feathered dinosaurs, Evolution of birds, having evolved from earlier Theropoda, theropods during the Late Jurassic epoch, and are the only dinosaur lineage known to have survived the Cretaceous–Paleogene extinction event approximately 66 mya. Dinosaurs can therefore be divided into avian dinosaurs—birds—and the extinct non-avian dinosaurs, which are all dinosaurs other than birds. Dinosaurs are varied from taxonomy (biology), taxonomic, ...
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Caenagnathoid
Oviraptorosaurs ("egg thief lizards") are a group of feathered maniraptoran dinosaurs from the Cretaceous Period of what are now Asia and North America. They are distinct for their characteristically short, beaked, parrot-like skulls, with or without bony crests atop the head. They ranged in size from ''Caudipteryx'', which was the size of a turkey, to the 8-meter-long, 1.4-ton ''Gigantoraptor''. The group (along with all maniraptoran dinosaurs) is close to the ancestry of birds. Some researchers such as Maryanska ''et al'' (2002) and Osmólska ''et al.'' (2004) have proposed that they may represent primitive flightless birds.Osmólska, Halszka, Currie, Philip J., Brasbold, Rinchen (2004) "The Dinosauria" Weishampel, Dodson, Osmólska. "Chapter 8 Oviraptorosauria" University of California Press. The most complete oviraptorosaur specimens have been found in Asia. The North American oviraptorosaur record is sparse.Varricchio, D. J. 2001. Late Cretaceous oviraptorosaur (Theropoda ...
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Elmisaurus
''Elmisaurus'' (meaning "foot sole lizard") is an extinct genus of caenagnathid dinosaur from the Late Cretaceous Nemegt Formation of Mongolia. It was a theropod belonging to the Oviraptorosauria. Discovery In 1970, a paleontological Polish-Mongolian expedition discovered two fragmentary specimens of a small theropod in the Ömnögovĭ province of Mongolia. The type species, ''Elmisaurus rarus'', was named and described by Halszka Osmólska in 1981. The generic name is derived from Mongol ' or ', "foot sole", as the type specimen consisted of a metatarsus. The specific name means "rare" in Latin. The holotype, ZPAL MgD-I/172, consists of a left metatarsus fused with the tarsalia. There are two paratypes: ZPAL MgD-I/98, consisting of a right hand and foot, and ZPAL MgD-I/20, the upper part of the left metatarsus of a larger individual.Osmólska, H. (1981). Coossified tarsometatarsi in theropod dinosaurs and their bearing on the problem of bird origins. ''Palaeontologica Polonic ...
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