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Diopsoidea Genera
The Diopsoidea are a small but diverse cosmopolitan superfamily of acalyptrate muscoids, especially prevalent in the tropics. Some flux exists in the family constituency of this group, with the Strongylophthalmyiidae and Tanypezidae formerly being in this group but now in the Nerioidea. Description As flies, Diopsoidea undergo complete metamorphosis with the four life stages of egg, larva, pupa and adult. The adult stage has three body segments (head, thorax and abdomen), three pairs of legs and one pair of wings. Some features that distinguish adult Diopsoidea from other flies are: a well-sclerotised face, antennae usually deflexed to strongly elbowed (if not, then either the fore or hind femur is entirely swollen), ocelli positioned near the vertex of the head, no katepisternal setae, wing veins R2+3 and R4+5 subparallel or slightly convergent apically, and the wing anal cell being comparable in size to the subcostal cell. References

Diopsoidea, Diptera superfamil ...
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Diopsidae
Stalk-eyed flies are insects of the fly family Diopsidae. The family is distinguished from most other flies by most members of the family possessing "eyestalks": projections from the sides of the head with the eyes at the end. Some fly species from other families such as Drosophilidae, Platystomatidae, Richardiidae, and Tephritidae have similar heads, but the unique character of the Diopsidae is that their Antenna (biology), antennae are located on the stalk, rather than in the middle of the head as in all other flies. Stalked eyes are present in all members of the subfamily Diopsinae, but are absent in the Centrioncinae, which retain unstalked eyes similar to those of other flies. The stalked eyes are usually sexually dimorphic, with eyestalks present but shorter in females. The stalk-eyed flies are up to a centimeter long, and they feed on both decaying plants and animals. Their unique morphology (biology), morphology has inspired research into how the attribute may have arisen ...
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Gobryidae
Gobryidae is a monotypic family of flies containing the genus ''Gobrya'', described by Francis Walker in 1860. They are relatively rare and found in Southeast Asia, Taiwan, Indonesia, Malaysia, the Philippines, and New Guinea. Gobryids are often metallic in coloration and their heads are wider than the rest of their bodies. Very little is known about the larval habits of these flies. McAlpine (1997) elevated the genus to family level. More research is needed to determine the best phylogenetic placement for ''Gobrya''. Genus and species The following are included in ''BioLib.cz''BioLib.cz: genus ''Gobrya'' Walker, 1860
(retrieved 21 March 2024)
and the

Megamerinidae
The Megamerinidae are a family of flies (Diptera) with about 11 species in three genera. They are small and are marked by an elongated, basally constricted abdomen. The family has been variously placed in the past within the superfamilies Diopsoidea, Nerioidea and more recently in Opomyzoidea but the evolutionary relationships remain unclear. Description For terms see Morphology of Diptera Dipteran morphology differs in some significant ways from the broader insect morphology, morphology of insects. The Diptera is a very large and diverse Order (biology), order of mostly small to medium-sized insects. They have prominent compound ey ...These are slender, medium-sized flies, with long abdomens. The hind femora are thickened, and bear two rows of spinules on the lower side. The costa is entire and the anal cell is elongated. They have no ocelli. Biology The biology of immature Megamerinidae is poorly known, but larvae have been recorded as predators living under bark or decay ...
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Nothybidae
The family Nothybidae contains only the genus ''Nothybus'', a group of colorful and elongated flies. The family has been recently revised. Taxonomy and description The morphology of Nothybidae is distinctive. The prothorax is elongated, with the front legs arising far back on the thorax, posterior to the anterior thoracic spiracles, making their heads appear to rest on a long neck. They are also distinguished by their narrow wing bases, patterned wings, rayed aristae, and the swollen subscutellum. Twelve nothybid species are described. First placed in Micropezidae, Frey recognized ''Nothybus'' as deserving a suprageneric rank, and Aczel also recognized the family Nothybidae. While this group is placed in the Diopsoidea, its evolutionary relationships have never been robustly tested in a modern phylogenetic framework with molecular or morphological data. It is one of the few fly families for which no DNA data are available on NCBI GenBank. Biology The biology of the Nothybidae ...
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Psilidae
The Psilidae are family of flies. Commonly called the rust flies, at least 38 species are in four genera. The carrot fly (''Chamaepsila rosae'') is a member of this group. They are found mainly in the Holarctic. Description For terms see Morphology of Diptera Dipteran morphology differs in some significant ways from the broader insect morphology, morphology of insects. The Diptera is a very large and diverse Order (biology), order of mostly small to medium-sized insects. They have prominent compound ey ... These are small or medium-sized (1.5 mm.-10 mm.) flies with slender bodies. They are yellow to reddish, brown or black in colour. The head is spherical with (relatively) small eyes and the face is often slanted backward. The antennae are small, with the third antennal segment conspicuously elongated. The arista has a short or long pubescence. The postvertical bristles are divergent or absent. Ocelli and ocellar bristles are present. Vibrissae are always absent. Up to ...
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Syringogastridae
''Syringogaster'' is a genus of small (4 to 6 mm) ant-mimicking flies with a petiolate abdomen, a long prothorax, a swollen and spiny hind femur, and reduced head size and large eyes. There are 20 described extant species and two species known from Miocene amber from the Dominican Republic. It is the only genus in the family Syringogastridae. Distribution The genus ''Syringogaster'' are found in tropical areas of the Neotropical region, with exception of the Antilles. Syringogastridae inhabit tropical forests. Biology Very little is known about the biology of Syringogastridae, and no larvae are known. Classification The descriptor Ezra Townsend Cresson placed his new genus in the family Psilidae The Psilidae are family of flies. Commonly called the rust flies, at least 38 species are in four genera. The carrot fly (''Chamaepsila rosae'') is a member of this group. They are found mainly in the Holarctic. Description For terms see Morpholo ... then in 1969 Angelo Pires ...
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Acalyptrate Muscoid
The Acalyptratae or Acalyptrata are a subsection of the Schizophora, which are a section of the order Diptera, the "true flies". In various contexts the Acalyptratae also are referred to informally as the acalyptrate muscoids, or acalyptrates, as opposed to the Calyptratae. All forms of the name refer to the lack of calypters in the members of this subsection of flies. An alternative name, Acalypterae is current, though in minority usage. It was first used by Pierre-Justin-Marie Macquart in 1835 for a section of his tribe Muscides; he used it to refer to all acalyptrates plus scathophagids and phorids, but excluding Conopidae. The confusing forms of the names stem from their first usage; ''Acalyptratae'' and ''Acalyptrata'' actually are adjectival forms in Neo-Latin. They were coined in the mid 19th century in contexts such as "Muscae Calyptratae and Acalyptratae" and "Diptera Acalyptrata", and the forms stuck. The Acalyptratae are a large assemblage, exhibiting very diverse h ...
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Strongylophthalmyiidae
The Strongylophthalmyiidae are a small Family (biology), family of about 80 species of slender, long-legged fly, flies, the majority of which occur in the Oriental and Australasian regions. They are divided into two genus, genera, the monotypic Southeast Asian genus ''Nartshukia'' Shatalkin, 1993 and ''Strongylophthalmyia'' Heller, 1902. The relationships of the group are obscure; formerly the genus ''Strongylophthalmyia'' was classified with the Psilidae, and some recent classifications place it within the Tanypezidae. Little is known of their biology, but many species seem to be associated with rotting bark. Species *Genus ''Strongylophthalmyia'' Heller, 1902 :*''Strongylophthalmyia angustipennis, S. angustipennis'' Axel Leonard Melander, Melander, 1920 :*''Strongylophthalmyia brunneipennis, S. brunneipennis'' (De Meijere, 1914) :*''Strongylophthalmyia caliginosa, S. caliginosa'' Iwasa, 1992 :*''Strongylophthalmyia coarctata, S. coarctata'' Friedrich Georg Hendel, Hendel, 1913 ...
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Tanypezidae
The Tanypezidae, known as the “stretched-foot flies”, are small family of acalyptrate Diptera (Schizophora, Brachycera). The 28 species are found mostly in the New World, divided between two genera: ''Tanypeza'' (2 species) is found in North America, with the type species (''T. longimana'' Fallén) extending into the Palaearctic, and ''Neotanypeza'' (26 species) is neotropical in distribution and includes one species known only from Dominican amber from 17 to 20 million years ago, ''N. dominicana'' Lonsdale & Apigian. This distribution contrasts that of its sister family, the Strongylophthalmyiidae, which is mostly East Asian in distribution. The family was recently treated by Lonsdale (2013), who redefined the family and its genera, synonymizing all other neotropical tanypezid genera in ''Neotanypeza''. Lonsdale (2014) also provided a full catalogue for the family. Species of Tanypezidae are relatively large and have semispherical heads and stout bodies that are perched ato ...
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Nerioidea
Nerioidea is a superfamily of Acalyptratae flies. Description As flies, Nerioidea undergo complete metamorphosis with the four life stages of egg, larva, pupa and adult. The adult stage has three body segments (head, thorax and abdomen), three pairs of legs and one pair of wings. Some features that distinguish adult Nerioidea from other flies are: a face that's usually weakly sclerotised (except in Fergusoninidae), antenna usually porrect or slightly deflexed (elbowed in Tanypezidae), wing veins R2+3 and R4+5 usually convergent, and the wing anal cell usually much smaller than the subcostal cell. Ecology Most Nerioidea are associated with dead and decaying organic matter such as dead wood, rotting fruit and bat dung. On the other hand, Fergusoninidae form galls in plants of family Myrtaceae, and some Micropezidae have larvae that are predatory or agricultural pests. Notable species * ''Glyphidops flavifrons ''Glyphidops flavifrons'' is a member of the Neriidae fami ...
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Metamorphosis
Metamorphosis is a biological process by which an animal physically develops including birth transformation or hatching, involving a conspicuous and relatively abrupt change in the animal's body structure through cell growth and differentiation. Some insects, jellyfish, fish, amphibians, mollusks, crustaceans, cnidarians, echinoderms, and tunicates undergo metamorphosis, which is often accompanied by a change of nutrition source or behavior. Animals can be divided into species that undergo complete metamorphosis (" holometaboly"), incomplete metamorphosis (" hemimetaboly"), or no metamorphosis (" ametaboly"). Generally organisms with a larval stage undergo metamorphosis, and during metamorphosis the organism loses larval characteristics. Etymology The word ''metamorphosis'' derives from Ancient Greek , "transformation, transforming", from ('), "after" and ('), "form". Hormonal control In insects, growth and metamorphosis are controlled by hormones synthesized by ...
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