Scarabaeidae
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Scarabaeidae
The family Scarabaeidae, as currently defined, consists of over 35,000 species of beetles worldwide; they are often called scarabs or scarab beetles. The classification of this family has undergone significant change. Several groups formerly treated as subfamilies have been elevated to family rank (e.g., Bolboceratidae, Geotrupidae, Glaresidae, Glaphyridae, Hybosoridae, Ochodaeidae, and Pleocomidae), and some reduced to lower ranks. The subfamilies listed in this article are in accordance with those in Catalog of Life (2023). Description Scarabs are stout-bodied beetles; most are brown or black in colour, but many, generally species that are diurnally active, have bright metallic colours, measuring between . The antenna (biology), antennae of most species superficially seem to be knobbed (capitate), but the several segments comprising the head of the antenna are, as a rule, lamellate: they extend laterally into plates called lamella (zoology), lamellae that they usually ...
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Cetoniinae
Flower chafers are a group of scarab beetles comprising the subfamily Cetoniinae. Many species are diurnal and visit flowers for pollen and nectar, or to browse on the petals. Some species also feed on fruit. The group is also called fruit and flower chafers, flower beetles and flower scarabs. There are around 4,000 species, many of them still undescribed. Ten tribes are presently recognized: Cetoniini, Cremastocheilini, Diplognathini, Goliathini, Gymnetini, Phaedimini, Schizorhinini, Stenotarsiini, Taenioderini, and Xiphoscelidini. The former tribes Trichiini and Valgini were elevated in rank to subfamily. The tribe Gymnetini is the biggest of the American tribes, and Goliathini contains the largest species, and is mainly found in the rainforest regions of Africa. Description Adult flower chafers are usually brightly coloured beetles, often metallic, and somewhat flattened in shape. The insertions of the antennae are visible from above, while the mandibles and lab ...
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Orphninae
Orphninae is a subfamily of beetles in the scarab beetle family, Scarabaeidae. There are two tribes in the family, the New World Aegidiini and the Old World The "Old World" () is a term for Afro-Eurasia coined by Europeans after 1493, when they became aware of the existence of the Americas. It is used to contrast the continents of Africa, Europe, and Asia in the Eastern Hemisphere, previously ... Orphnini.Jameson, M. L. and F. C. OcampoOrphninae.Generic Guide to New World Scarab Beetles. University of Nebraska State Museum. 2005 Version. They are mostly tropical beetles.Frolov, A. (2013)''Stenosternus'' Karsch, a possible link between Neotropical and Afrotropical Orphninae (Coleoptera, Scarabaeidae).''ZooKeys'' 335 33-46. As of 2013 there are about 195 species in the subfamily. They are classified in 15 genera: 4 in the tribe Aegidiini and 11 in the Orphnini. In addition, the genus ''Stenosternus'' is provisionally placed in the Aegidiini. Taxonomy Taxa include:Frolov, ...
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Melolonthinae
Melolonthinae is a subfamily of the Scarabaeidae, scarab beetles (family (biology), family Scarabaeidae). It is a very diverse group; distributed over most of the world, it contains over 11,000 species in over 750 genera. Some authors include the scarab subfamilies Euchirinae and Pachypodinae as tribe (biology), tribes in the Melolonthinae. Unlike some of their relatives, their Morphology (biology), habitus is usually not bizarre. They resemble the Rutelinae in being fairly plesiomorphic in outward appearance. Like in many Scarabaeidae, males have large fingered antenna (biology), antennae, while those of the females are smaller and somewhat knobby. In the Melolonthinae, this sexual dimorphism is particularly pronounced. Many species have striking – though rarely brilliant or iridescent – hues and bold patterns of hairs. Being often quite sizeable and swarming in numbers at certain times, for example the ''Amphimallon'', ''Phyllophaga'' and ''Polyphylla'' "June beetle ...
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Aphodiinae
Aphodiinae is a subfamily of the scarab beetle family, Scarabaeidae. Members of this subfamily are known commonly as the small dung beetles and many, but not all, are dung beetles.Skelley, P. EAphodiinae. Generic Guide to New World Scarab Beetles. University of Nebraska State Museum. 2008 Version. These beetles are found worldwide. These beetles are small scarab beetles, most less than 8 millimeters long. Many have small mandibles that are covered by a widened clypeus, the exoskeleton plate above the mouth. The feet are clawed. This is a diverse subfamily with varied life strategies and habitat types. Many species are dung beetles, which collect and feed on animal dung. Other species are detritivores or saprophages, which feed on dead matter, and some are predatory. Some are known as inquilines, living in ant or termite nests, and some are sand-dwelling beetles. A survey of South American aphodiines found them in diverse habitat types including temperate rainforests, high-ele ...
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Rutelinae
Rutelinae or shining leaf chafers is a subfamily of the scarab beetles (family Scarabaeidae). It is a very diverse group; distributed over most of the world, it contains some 200 genera with over 4,000 described species in 7 tribes. Several taxa have yet to be described. A few recent classifications include the tribe Hopliini, but this is not generally accepted. Description Unlike some of their relatives, their habitus is usually lacking in ornamentation, such as horns. They resemble the Melolonthinae in being fairly plesiomorphic in outward appearance. Many species have brilliant or iridescent hues, however, such as the genus '' Chrysina'', and a number of species are serious pests (e.g., the Japanese beetle). Behavior Feeding Adult Rutelinae feed on leaves, flowers, and flower parts. Larvae feed on decaying wood, compost or roots. Tribes * Adoretini * Alvarengiini * Anatistini (= Spodochlamyini) * Anomalini * Anoplognathini * Geniatini * Rutelini Additionally t ...
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Aclopinae
Aclopinae is a subfamily of Scarabaeidae or scarab beetles in the superfamily Scarabaeoidea Scarabaeoidea is a superfamily of beetles, the only subgroup of the infraorder Scarabaeiformia. Around 35,000 species are placed in this superfamily and some 200 new species are described each year. Some of its constituent families are undergo .... Distribution The subfamily is found in northern Australia, Borneo and southern South America (Brazil and Argentina). Genera These 11 genera belong to the subfamily Aclopinae. ; Tribe Aclopini Blanchard, 1850 : '' Aclopus'' Erichson, 1835 (Neotropics) : '' Desertaclopus'' Ocampo & Mondaca, 2012 : '' Gracilaclopus'' Ocampo & Mondaca, 2012 : †'' Cretaclopus'' Nikolajev, 2004 : †'' Juraclopus'' Nikolajev, 2005 (Upper Jurasic) : †'' Prophaenognatha'' Bai, Ren & Yang, 2011 ; Tribe Holcorobeini Nikolajev, 1992 (Mesozoic) : †'' Antemnacrassa'' Gomez Pallerola, 1979 : †'' Holcorobeus'' Nikritin, 1977 (Lower Cretaceous) : †'' Mongolrobeu ...
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Aegialiinae
Aegialiinae is a small subfamily of the family Scarabaeidae (scarab beetles).Aegialiinae at Zipcodezoo
Historically the group has been treated as a tribe within a broad definition of the subfamily Aphodiinae.


Genera

The following genera are members of the subfamily Aegialiinae. * Tribe Aegialiini ** ''Aegialia'' (Palaearctic, Nearctic, Neotropical) ** ''Caelius (beetle), Caelius'' (Nearctic, Palaearctic) ** ''Micraegialia'' (the Nearctic) ** ''Psammoporus'' (Palaearctic, Nearctic) ** ''Rhysothorax'' (Palaearctic, Nearctic) ** ''Silluvia'' (Palaearctic, Oriental) ** † ''Archeopsammoporus'' (Palaearctic, Fossil) ** † ''Cretaegialia'' (Fossil) * Tribe Saprinianini ** ''Amerisaprus'' (Neotropical) ** ''Argeremazus'' ...
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Allidiostomatinae
Allidiostomatinae is a subfamily of beetles in the scarab beetle family, Scarabaeidae. It is distributed in southern South America. Of the eleven species, seven are endemism, endemic to Argentina. Others can also be found in Chile and Peru.Ocampo, F. C. and J. Colby. (2009)''Parallidiostoma tricornum'' Ocampo and Colby, a new genus and species of Allidiostomatinae from Peru (Coleoptera: Scarabaeidae).''Zootaxa'' 2287 64-68. Little is known about the biology of these beetles.Jameson, M. L. and F. C. Ocampo Generic Guide to New World Scarab Beetles. University of Nebraska State Museum. 2005 Version. The subfamily was made up of the single genus ''Allidiostoma'' until 2009, when a second was erected for the new species ''Parallidiostoma tricornum''. Genera and species include: ; Genus ''Allidiostoma'' Arrow, 1940 : ''Allidiostoma bosqui'' Gutiérrez, 1946 - Chile : ''Allidiostoma halffteri'' Martinez, 1956 - Argentina : ''Allidiostoma hirta'' (Ohaus, 1910) - Argentina : ''Allid ...
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Scarabaeinae
The scarab beetle subfamily Scarabaeinae consists of species collectively called true dung beetles (there are also dung beetles in other subfamilies and families). Most of the beetles of this subfamily feed exclusively on dung. However, some may feed on decomposing matter including carrion, decaying fruits and fungi. Dung beetles can be placed into three structural guilds based on their method of dung processing namely rollers (telecoprids), dwellers (endocoprids) and tunnelers (paracoprids). Dung removal and burial by dung beetles result in ecological benefits such as soil aeration and fertilization; improved nutrient cycling and uptake by plants, increase in pasture quality, biological control of pest flies and intestinal parasites and secondary seed dispersal. Well-known members include the genera '' Scarabaeus'' and '' Sisyphus'', and '' Phanaeus vindex''. Description Adult dung beetles have modified mouth parts which are adapted to feeding on dung. The clypeus is expan ...
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Dynamopodinae
Dynamopodinae is a very small subfamily of Scarabaeidae or scarab beetles, with only seven species in a two genera.Google Books
John L. Capinera Encyclopedia of Entomology — 2nd Edition. 3279 pp.


Genera and Species

These genera and species belong to the subfamily Dynamopodinae: ; Genus ''Adraria'' Villiers, 1956 : ''Adraria monodi'' Villiers, 1956 - Mauretania ; Genus ''Orubesa'' Reitter, 1895 : ''Orubesa ata'' (Semenov & Medvedev, 1929) - Iran : ''Orubesa athleta'' (Semenov, 1895) - Russia, India, Central Asia : ''Orubesa gladiator'' (Balthasar, 1968) - Afghanistan : ''Orubesa luctator'' (Semenov & Medvedev, 1929) - Iraq, Iran, P ...
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Termitotroginae
Termitotroginae is a monotypic subfamily of the family Scarabaeidae, the scarab beetles. The only genus in the subfamily is ''Termititrox''. A second genus, ''Aphodiocopris'', Arrow, 1920, has been synonym (taxonomy), synonymised with ''Termitotrox''. All known members of this subfamily are tiny, blind and flightless, and are Termitophile, termitophiles, dwelling within the fungal gardens of colonies of species of termite in Africa or tropical Asia. Termitotrogins are characterised by having no eyes and no wings, and their pronotum and elytra have a distinctive patterning of ribs and grooves. The function of the beetles in the termite colony is unclear but it seems that they are likely to be obligatory termitophiles and somehow play an important role in the nest environment of their fungus-growing hosts. Species The following species are included in the genus ''Termitotrox'': *''Termitotrox ancoroides'' (Petrovitz, 1956) – Eastern Democratic Republic of the Congo *''Termitotr ...
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Oncerinae
Oncerinae is a small subfamily of scarab beetles in the family Scarabaeidae. There are only two genera Oncerinae, each with a single species. Oncerinae has been sometimes classified as a tribe of the subfamily Melolonthinae, "Oncerini". Genera These two genera belong to the subfamily Oncerinae: ; Genus '' Nefoncerus'' Saylor, 1938 - Neotropics The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropical terrestrial ecoregions of the Americas and the entire South American temperate zone. Definition In biogeog ... : '' Nefoncerus convergens'' (Horn, 1894) ; Genus '' Oncerus'' LeConte, 1856 - North America : '' Oncerus floralis'' LeConte, 1856 References Scarabaeidae Articles created by Qbugbot {{scarabaeidae-stub ...
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