Nematocera
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Nematocera
The Nematocera (the name meaning "thread-horns") are a suborder of elongated fly, flies with thin, segmented antenna (biology), antennae and mostly aquatic larvae. This group is paraphyletic and contains all flies except for species from suborder Brachycera (the name meaning "short-horns"), which includes more commonly known species such as the housefly or the Drosophila_melanogaster, common fruit fly. The equivalent clade to Nematocera is the whole Diptera, with Brachycera as a subclade. Families in Nematocera include mosquitoes, crane fly, crane flies, gnats, Black fly, black flies, and multiple families commonly known as midges. The Nematocera typically have fairly long, fine, finely-jointed antennae. In many species, such as most mosquitoes, the female antennae are more or less threadlike, but the males have spectacularly wikt:plumose, plumose antennae. The larvae of most families of Nematocera are aquatic, either free-swimming, rock-dwelling, plant-dwelling, or wikt:luticolo ...
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Axymyiomorpha
The Nematoceran family Axymyiidae is the sole member of the infraorder Axymyiomorpha, though it is often included within the infraorder Bibionomorpha in older classifications. It is known from only nine species in four genera, plus eight fossil species. Family characteristics The Axymyiidae have the general appearance of the Bibionidae Bibionidae (March flies) is a Family (biology), family of Fly, flies (Fly, Diptera) containing approximately 650–700 species worldwide. Adults are nectar feeders and emerge in numbers in spring. Because of the likelihood of adult flies being f .... Unlike bibionids, axymyiids have four branches of the radial vein, Bibionidae have two or three. The head is rounded. The eyes of the male are holoptic for a considerable distance and divided into a larger dorsal part consisting of large facets and (separated by a groove) a smaller ventral part of smaller facets. The eyes of the female are separated by a broad frons and consist of separated fac ...
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Perissommatomorpha
The Perissommatidae are a family of flies (Diptera) that was proposed in 1962 by Donald Colless based on the species ''Perissomma fusca'' from Australia. The family now includes five extant species within the single genus ''Perissomma'', four from Australia and one from Chile. The Perissommatidae are unusual as they appear to have four compound eyes. They have a small slender body less than 2 mm in length. Their wings are large in comparison to their bodies and subsequently their flight is weak. Preferring high-altitude forest environments, adults only fly in the winter. The larvae live in decaying leaf litter in wet sclerophyll or cool rain forests. Some species are suspected to be associated with fungi. In the case of ''Perissomma macalpinei'', numbers of adults have been observed congregating in clumps of foliage and rising in short, zigzag flights in the sunlight above the foliage for short periods before descending. Fossils of the family are known from the Middle Jurassi ...
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Culicomorpha
The Culicomorpha are an infraorder of Nematocera, including mosquitoes, black flies, and several extant and extinct families of insects. They originated 176 million years ago, in the Triassic period. There are phylogenetic patterns that are used to interpret bionomic features such as differences in the nature of blood-feeding by adult females, daytime or nighttime feeding by adult females, and occurrence of immature stages in aquatic habitats. Most adult, females lay their eggs on bodies of water. Some are restricted to very clean waters, but others can tolerate highly polluted environments. Many adults transmit parasites or diseases that can be debilitating or fatal to humans, such as malaria and West Nile virus. Biting midges can transmit an extensive range of pathogens of veterinary importance, including Akabane virus, bovine ephemeral fever virus, Schmallenberg virus, African horse sickness virus, epizootic haemorrhagic disease virus, and bluetongue virus. Classificat ...
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Nymphomyiomorpha
The Nymphomyiidae are a family of tiny (2 mm) slender, delicate flies (Diptera). Larvae are found among aquatic mosses in small, rapid streams in northern regions of the world, including northeastern North America, Japan, the Himalayas, and eastern Russia. Around a dozen extant species are known, with two fossil species found in amber, extending back to the Mid Cretaceous. Under an alternative classification, they are considered the only living representatives of a separate, suborder called Archidiptera (or Archaediptera) which includes several Triassic fossil members. The family has characteristics associated with the Nematocera as well as the Brachycera. The antennae are shortened as in the Brachycera and these flies are long, having a snout with vestigeal mouthparts, non-differentiated abdominal segments with large cerci. The wings are narrow and hair-fringed and have very weak venation. They are known to form cloud-like swarms in summer and the short-lived non-feeding ad ...
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Midge
A midge is any small fly, including species in several family (biology), families of non-mosquito nematoceran Diptera. Midges are found (seasonally or otherwise) on practically every land area outside permanently arid deserts and the frigid zones. Some midges, such as many Phlebotominae (sand fly) and Simuliidae (black fly), are vectors of various diseases. Many others play useful roles as prey for insectivores, such as various Wallum Sedge Frog, frogs and Hirundinidae, swallows. Others are important as detritivores, and form part of various nutrient cycles. The habits of midges vary greatly from species to species, though within any particular family, midges commonly have similar ecological roles. Examples of families that include species of midges include: * Blephariceridae, net-winged midges * Cecidomyiidae, gall midges * Ceratopogonidae, biting midges (also known as no-see-ums or punkies in North America and sandflies in Australia) * Chaoboridae, phantom midges * Chiron ...
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Crane Fly
A crane fly is any member of the dipteran superfamily Tipuloidea, which contains the living families Cylindrotomidae, Limoniidae, Pediciidae and Tipulidae, as well as several extinct families. "Winter crane flies", members of the family Trichoceridae, are sufficiently different from the typical crane flies of Tipuloidea to be excluded from the superfamily Tipuloidea, and are placed as their sister group within Tipulomorpha. Two other families of flies, the phantom crane flies ( Ptychopteridae) and primitive crane flies ( Tanyderidae), have similar common names due their similar appearance, but they are not closely related to true crane flies. The classification of crane flies has been varied in the past, with some or all of these families treated as subfamilies, but the following classification is currently accepted. Species counts are approximate, and vary over time. ; Infraorder Tipulomorpha : Superfamily Tipuloidea (Typical Crane Flies) :: Family Cylindrotomidae (Cy ...
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Mycetophilidae
Mycetophilidae is a family of small flies, forming the bulk of those species known as fungus gnats. About 3000 described species are placed in 150 genera, but the true number of species is undoubtedly much higher. They are generally found in the damp habitats favoured by their host fungi and sometimes form dense swarms. Adults of this family can usually be separated from other small flies by the strongly humped thorax, well-developed coxae, and often spinose legs, but identification within the family between genera and species generally requires close study of microscopic features such as subtle differences in wing venation and variation in chaetotaxy and genitalia. The terrestrial larvae usually feed on fungi, especially the fruiting bodies, but also spores and hyphae, but some species have been recorded on mosses and liverworts. The larvae of some species, while still being associated with fungi, are at least partly predatory. Some species are attracted to the fungus smell o ...
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Deuterophlebiomorpha
''Deuterophlebia'', also known as mountain midges are a genus of flies that are the sole living members of the small family Deuterophlebiidae. Adults have broad, fan-shaped wings, and males have extremely long antennae which they employ when contesting territories over running water, waiting for females to hatch. Larvae occur in swiftly flowing streams and are easily recognized by their forked antennae and the prolegs on the abdomen. One classification places this family in its own infraorder Deuterophlebiomorpha, but this has not gained wide acceptance. A recent phylogeny of the entire order Diptera places them as the sister group to all other flies. Fossil members of the family are known from the mid-Cretaceous Burmese amber of Myanmar, dating to around 100 million years ago. Species Species of the genus are known from Afghanistan, Kashmir, Northeast India, China, Japan, Korea, Eastern Siberia, and western North America from Alaska to California to Colorado. Catalogue of ...
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Bibionomorpha
The Bibionomorpha are an infraorder of the suborder Nematocera. One of its constituent families, the Anisopodidae, is the presumed sister taxon to the entire suborder Brachycera. Several of the remaining families in the infraorder (those shown without common names) are former subfamilies of the Mycetophilidae, which has been recently subdivided. The family Axymyiidae has recently been removed from the Bibionomorpha to its own infraorder Axymyiomorpha. Most representatives of the Bibionomorpha are saprophages or fungivores as larvae with the Cecidomyiidae being predominantly gall-formers. Some sciarids are common indoor pests, developing large populations in potting soil that has become moldy from overwatering. The larvae of the Bibionidae sometimes migrate in large, snake-like masses to minimize dehydration while seeking a new feeding site. Superfamilies and families *Superfamily Anisopodoidea (Triassic - Recent) **Anisopodidae (syn † Eopleciidae) - wood gnats **† Antef ...
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Tipulidae
Tipulidae is a family of large Crane fly, crane flies in the order Diptera. There are more than 30 genera and 4,200 described species in Tipulidae, common and widespread throughout the world. A crane fly can be identified as a member of Tipulidae by its maxillary palps, which is the pair of appendages that hang down from the front of its head. If the fourth segment (the furthest from the body) of the maxillary palp is longer than the other three combined, then it is likely to be a member of Tipulidae. There are also usually 13 segments in the antennae of large crane flies, compared to 14 or 16 in the common Limoniidae, limoniid crane flies. The oldest fossils that can be assigned confidently to Tipulidae ''sensu stricto'' are those of the genus ''Tipunia,'' which date to the Late Jurassic. Genera These 39 genera belong to the family Tipulidae: * ''Acracantha'' Skuse, 1890 * ''Angarotipula'' Savchenko, 1961 * ''Austrotipula'' Alexander, 1920 * ''Brachypremna'' Osten Sacken, 1 ...
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Ptychopteromorpha
Ptychopteromorpha is a taxonomic group within the suborder Nematocera The Nematocera (the name meaning "thread-horns") are a suborder of elongated fly, flies with thin, segmented antenna (biology), antennae and mostly aquatic larvae. This group is paraphyletic and contains all flies except for species from suborder ... consisting of two uncommon families. In older classifications, these families were included within the infraorder Tipulomorpha, based on superficial similarities (e.g., slender bodies and long legs). The inclusion of the families Tanyderidae and Ptychopteridae was based on the foldability of the last tarsomere in males. Molecular studies show no close relationship between the Tanyderidae and the Ptychopteridae, and support for this grouping is limited. References External links The Tree of Life Project Insect infraorders {{Ptychopteromorpha-stub ...
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Tipulomorpha
The Tipulomorpha are an infraorder of Nematocera, containing the crane flies, a very large group, and allied families. Taxonomy The Tipulomorpha comprise five extant families. *Tipulomorpha **Tipuloidea (typical crane flies) *** Cylindrotomidae (long-bodied crane flies) *** Pediciidae (hairy-eyed crane flies) ***Limoniidae (limoniid crane flies) – probably paraphyletic ***Tipulidae (large crane flies) ** Trichoceroidea (winter crane flies) *** Trichoceridae Extinct superfamilies One recent classification based largely on fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...s splits this group into a series of extinct superfamilies (below), and includes members of other infraorders, but this has not gained wide acceptance. *Superfamily Eopolyneuroidea **Family Eop ...
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