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The Perissommatidae are a family of
flies Flies are insects of the Order (biology), order Diptera, the name being derived from the Ancient Greek, Greek δι- ''di-'' "two", and πτερόν ''pteron'' "wing". Insects of this order use only a single pair of wings to fly, the hindwing ...
(Diptera) that was proposed in 1962 by Donald Colless based on the species ''Perissomma fusca'' from Australia. The family now includes five extant species within the single genus ''Perissomma'', four from Australia and one from
Chile Chile, officially the Republic of Chile, is a country in the western part of South America. It is the southernmost country in the world, and the closest to Antarctica, occupying a long and narrow strip of land between the Andes to the eas ...
. The Perissommatidae are unusual as they appear to have four compound eyes. They have a small slender body less than 2 mm in length. Their wings are large in comparison to their bodies and subsequently their flight is weak. Preferring high-altitude forest environments, adults only fly in the winter. The larvae live in decaying leaf litter in wet sclerophyll or cool rain forests. Some species are suspected to be associated with fungi. In the case of ''Perissomma macalpinei'', numbers of adults have been observed congregating in clumps of foliage and rising in short, zigzag flights in the sunlight above the foliage for short periods before descending. Fossils of the family are known from the Middle Jurassic to Early Cretaceous of Eastern and Northern Asia.


Extinct genera

After * †'' Limnorhyphus'' Hong 1983,
Haifanggou Formation The Haifanggou Formation (), also known as the Jiulongshan Formation (), is a fossil-bearing rock deposit located near Daohugou () village of Ningcheng County, in Inner Mongolia, northeastern China. The formation consists of coarse conglomerate ...
, China, Middle Jurassic (
Callovian In the geologic timescale, the Callovian is an age and stage in the Middle Jurassic, lasting between 166.1 ± 4.0 Ma (million years ago) and 163.5 ± 4.0 Ma. It is the last stage of the Middle Jurassic, following the Bathonian and preceding the ...
) * Subfamily Perissommatinae Colless 1962 ** †'' Collessomma'' Lukashevich and Blagoderov 2020 Khasurty locality, Russia, Dzun-Bain Formation, Mongolia, Early Cretaceous (
Aptian The Aptian is an age in the geologic timescale or a stage in the stratigraphic column. It is a subdivision of the Early or Lower Cretaceous Epoch or Series and encompasses the time from 121.4 ± 1.0 Ma to 113.0 ± 1.0 Ma (million years ag ...
) ** †'' Gurvaniella'' Kovalev 1986 Gurvan-Eren Formation, Mongolia, Aptian ** †'' Palaeoperissomma'' Kovalev 1985
Itat Formation The Itat Formation ( Russian: итатская свита) is a geologic formation in western Siberia. It was deposited in the Bajocian to Bathonian ages of the Middle Jurassic. Dinosaur remains are among the fossils that have been recovered from ...
, Russia, Middle Jurassic (
Bathonian In the geologic timescale the Bathonian is an age and stage of the Middle Jurassic. It lasted from approximately 168.3 Ma to around 166.1 Ma (million years ago). The Bathonian Age succeeds the Bajocian Age and precedes the Callovian Age. St ...
), Shar Teeg, Mongolia, Late Jurassic (
Tithonian In the geological timescale, the Tithonian is the latest age of the Late Jurassic Epoch and the uppermost stage of the Upper Jurassic Series. It spans the time between 152.1 ± 4 Ma and 145.0 ± 4 Ma (million years ago). It is preceded by t ...
) Turga Formation, Russia, Aptian * †subfamily Rasnicynommatinae Lukashevich 2011 ** †'' Rasnicynomma'' Lukashevich 2011 Shar Teeg, Mongolia, Tithonian * †subfamily Perissordinae Lukashevich et al. 2006 ** †'' Perissordes'' Lukashevich et al. 2006
Daohugou The Haifanggou Formation (), also known as the Jiulongshan Formation (), is a fossil-bearing rock deposit located near Daohugou () village of Ningcheng County, in Inner Mongolia, northeastern China. The formation consists of coarse conglomerate ...
, China, Callovian


References


External links

*Short account By Evenhuis, N. L

Nematocera families {{Nematocera-stub