Mormyridae
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Mormyridae
The Mormyridae, sometimes called "elephantfish" (more properly freshwater elephantfish), are a superfamily of weakly electric fish in the order Osteoglossiformes native to Africa. It is by far the largest family in the order, with around 200 species. Members of the family can be popular, if challenging, aquarium species. These fish have a large brain size and unusually high intelligence. They are not to be confused with the marine and brackish-water callorhinchid elephantfish (family Callorhinchidae) of Southern Hemisphere oceans. Description and biology The elephantfish are a diverse family, with a wide range of different sizes and shapes. The smallest are just in adult length, while the largest reach up to . They do, however, have a number of unique features in common. Firstly, their cerebellum is greatly enlarged, enabling them to interpret complex bio-electrical signals, and to the large size of the valve. Secondly, an auditory vesicle (a small bladder) is present insid ...
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Mormyromast Diagram
The Mormyroidea (Synonym (taxonomy), synonymy: Mormyriformes) are a superfamily (formerly an order) of fresh water fishes Endemism, endemic to Africa that, together with the families ''Mooneye, Hiodontidae, Osteoglossidae, Pantodontidae'' and ''Notopteridae'', represents one of the main groups of living Osteoglossiformes. They stand out for their Electroreception and electrogenesis, use of weak electric fields, which they use to orient themselves, reproduce, feed, and communicate. There is no consensus regarding its superior Taxonomy (biology), biological classification as some experts state that it belongs to the Order (biology), suborder ''Osteoglossoidei'', while others to the ''Notopteroidei''. In either case, the mormyriformes include the Gymnarchus, gymnarchids and Mormyridae, mormyrids and represent the largest superfamily within the order ''Osteoglossiformes'' with about two hundred and thirty-three subordinate Taxon, taxa that are Species distribution, distributed across ...
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Electric Organ (biology)
An Torpediniformes.html" ;"title="electric ray (Torpediniformes">electric ray (Torpediniformes) showing location of paired electric organs in the head, and electrocytes stacked within it In biology, the electric organ is an organ (biology), organ that an electric fish uses to create an electric field. Electric organs are derived from modified muscle or in some cases nerve tissue, called electrocytes, and have evolved at least six times among the Elasmobranchii, elasmobranchs and teleosts. These fish use their electric discharges for navigation, communication, mating, defence, and in strongly electric fish also for the incapacitation of prey. The electric organs of two strongly electric fish, the torpedo ray and the electric eel, were first studied in the 1770s by John Walsh, Hugh Williamson, and John Hunter. Charles Darwin used them as an instance of convergent evolution in his 1859 ''On the Origin of Species''. Modern study began with Hans Lissmann's 1951 study of electr ...
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Gymnarchus
The African knifefish, ''Gymnarchus niloticus'' – also called the ''aba aba'' – is an electric fish, living at the bottoms of rivers and lakes. It is the only species in the genus ''Gymnarchus'' and the family Gymnarchidae, within the order Osteoglossiformes. It is a long slender fish with no pelvic or anal fins, and a tail fin shaped like a rat's tail. It swims using its elongated dorsal fin, allowing it to keep its body straight while it moves. This in turn enables it to produce a steady but weak electric field, which it uses to locate its prey. It is large for a river fish; adults can reach 1.6 m (5.2 ft) in length and 19 kg (42 lb) in weight. In 1950, Hans Lissmann noticed that the fish could swim equally well forwards or backwards, clearly relying on a sense other than vision. He demonstrated that it could locate prey by electroreception, making it the first fish known to have this ability. The fish is considered good to eat in West Africa, wh ...
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Gnathonemus Petersii
Peters's elephant-nose fish (''Gnathonemus petersii'') is an African freshwater elephantfish in the genus '' Gnathonemus''. Other names in English include elephantnose fish, long-nosed elephant fish, and Ubangi mormyrid, after the Ubangi River. The Latin name is probably for the German naturalist Wilhelm Peters. The fish uses electrolocation to find prey, and has the largest brain-to-body oxygen use ratio of all known vertebrates (around 0.6).Nilsson G (1996"Brain and body oxygen requirements of Gnathonemus petersii, a fish with an exceptionally large brain"''Journal of Experimental Biology'', 199(3): 603–607Download/ref> Description Peters's elephantnose fish is native to the rivers of West and Central Africa, in particular the lower Niger River basin, the Ogun River basin and the upper Chari River. It prefers muddy, slowly moving rivers and pools with cover such as submerged branches. The fish is a dark brown to black in colour, laterally compressed (averaging ), with a r ...
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Electric Fish
An electric fish is any fish that can Bioelectrogenesis, generate electric fields, whether to sense things around them, for defence, or to stun prey. Most fish able to produce shocks are also electroreceptive, meaning that they can sense electric fields. The only exception is the Uranoscopidae, stargazer family (Uranoscopidae). Electric fish, although a small minority of all fishes, include both oceanic and freshwater species, and both cartilaginous and bony fishes. Electric fish produce their electrical fields from an Electric organ (biology), electric organ. This is made up of electrocytes, modified muscle or nerve cells, specialized for producing strong electric fields, used to locate prey, for Anti-predator adaptation, defence against predators, and for Signalling theory, signalling, such as in courtship. Electric organ discharges are two types, pulse and wave, and vary both by species and by function. Electric fish have evolved many specialised behaviours. The predatory Cla ...
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Osteoglossiformes
Osteoglossiformes , meaning "bony tongues" in Ancient Greek, is a relatively primitive order of ray-finned fish that contains two sub-orders, the Osteoglossoidei and the Notopteroidei. All of at least 245 living species inhabit freshwater. They are found in South America, Africa, Australia and southern Asia, having first evolved in Gondwana before that continent broke up. In 2008, several new species of marine osteoglossiforms were described from the Danish Eocene Fur Formation, dramatically increasing the diversity of this group. This implies that the Osteoglossomorpha is not a primary freshwater fish group with the osteoglossiforms having a typical Gondwana distribution. The Gymnarchidae (the only species being '' Gymnarchus niloticus'', the African knifefish) and the Mormyridae are weakly electric fish able to sense their prey using electric fields. The mooneyes (Hiodontidae) are often classified here, but may also be placed in a separate order, Hiodontiformes. Members ...
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Knollenorgan
A Knollenorgan is an electroreceptor in the skin of weakly electric fish of the family Mormyridae (Elephantfish) from Africa. The structure was first described by Viktor Franz (1921), a German anatomist unaware of its function. They are named after "Knolle", German for "tuberous root" which describes their structure. Structure and function Knollenorgans contain modified Epithelium, epithelial cells that act as Transduction (physiology), sensory transducers for electric fields. Besides these, there are supporting cells and a sensory neuron. The neuron projects to the fish's brain, specifically to the nucleus of the electrosensory lateral line lobe (nELL) of the medulla oblongata, medulla via the posterior branch of the lateral line nerve. The organs are embedded in the thickened Epidermis (skin), epidermis. The receptor cells lie buried in the deeper layers of the epidermis, where they expand into a pocket in the superficial layers of the Dermis, corium. The sense organ is surro ...
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Electric Discharge
In electromagnetism, an electric discharge is the release and transmission of electricity in an applied electric field through a medium such as a gas (i.e., an outgoing flow of electric current through a non-metal medium).American Geophysical Union, National Research Council (U.S.). Geophysics Study Committee (1986) ''The earth's electrical environment''. National Academy Press, Washington, DC, p. 263. Applications The properties and effects of electric discharges are useful over a wide range of magnitudes. Tiny pulses of current are used to detect ionizing radiation in a Geiger–Müller tube. A low steady current can illustrate the gas spectrum in a gas-filled tube. A neon lamp is an example of a gas-discharge lamp, useful both for illumination and as a voltage regulator. A flashtube generates a short pulse of intense light useful for photography by sending a heavy current through a gas arc discharge. Corona discharges are used in photocopiers. Electric discharges can convey ...
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Conductivity (electrolytic)
Conductivity or specific conductance of an electrolyte solution is a measure of its ability to conduct electricity. The SI unit of conductivity is siemens per meter (S/m). Conductivity measurements are used routinely in many industrial and environmental applications as a fast, inexpensive and reliable way of measuring the ionic content in a solution. For example, the measurement of product conductivity is a typical way to monitor and continuously trend the performance of water purification systems. In many cases, conductivity is linked directly to the total dissolved solids (TDS). High-quality deionized water has a conductivity of : \kappa = 0.05501 \pm 0.0001 μS/cm at 25 °C. This corresponds to a specific resistivity of : \rho = 18.18 \pm 0.03 MΩ⋅cm. The preparation of salt solutions often takes place in unsealed beakers. In this case the conductivity of purified water often is 10 to 20 times higher. A discussion can be found below. Typical drinking water is ...
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Ampullae Of Lorenzini
Ampullae of Lorenzini (: ''ampulla'') are electroreceptors, sense organs able to detect electric fields. They form a network of mucus-filled pores in the skin of Chondrichthyes, cartilaginous fish (sharks, Ray (fish), rays, and chimaeras) and of basal Osteichthyes, bony fishes such as reedfish, sturgeon, and lungfish. They are associated with and evolved from the mechanosensory lateral line organs of early vertebrates. Most bony fishes and terrestrial vertebrates have lost their ampullae of Lorenzini. History Ampullae were initially described by Marcello Malpighi and later given an exact description by the Italian physician and ichthyologist Stefano Lorenzini in 1679, though their function was unknown. Electrophysiology, Electrophysiological experiments in the 20th century suggested a sensibility to temperature, mechanical pressure, and possibly salinity. In 1960 the ampullae were identified as specialized receptor organs for sensing electric fields. One of the first descriptio ...
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Electric Field
An electric field (sometimes called E-field) is a field (physics), physical field that surrounds electrically charged particles such as electrons. In classical electromagnetism, the electric field of a single charge (or group of charges) describes their capacity to exert attractive or repulsive forces on another charged object. Charged particles exert attractive forces on each other when the sign of their charges are opposite, one being positive while the other is negative, and repel each other when the signs of the charges are the same. Because these forces are exerted mutually, two charges must be present for the forces to take place. These forces are described by Coulomb's law, which says that the greater the magnitude of the charges, the greater the force, and the greater the distance between them, the weaker the force. Informally, the greater the charge of an object, the stronger its electric field. Similarly, an electric field is stronger nearer charged objects and weaker f ...
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Flagellum
A flagellum (; : flagella) (Latin for 'whip' or 'scourge') is a hair-like appendage that protrudes from certain plant and animal sperm cells, from fungal spores ( zoospores), and from a wide range of microorganisms to provide motility. Many protists with flagella are known as flagellates. A microorganism may have from one to many flagella. A gram-negative bacterium '' Helicobacter pylori'', for example, uses its flagella to propel itself through the stomach to reach the mucous lining where it may colonise the epithelium and potentially cause gastritis, and ulcers – a risk factor for stomach cancer. In some swarming bacteria, the flagellum can also function as a sensory organelle, being sensitive to wetness outside the cell. Across the three domains of Bacteria, Archaea, and Eukaryota, the flagellum has a different structure, protein composition, and mechanism of propulsion but shares the same function of providing motility. The Latin word means " whip" to describe its ...
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