Hymenocarina
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Hymenocarina
Hymenocarina is an Order (biology), order of extinct marine arthropods known from the Cambrian. They possess bivalved carapaces, typically with exposed posteriors. Members of the group are morphologically diverse and had a variety of ecologies, including as filter feeders and as predators. Recent research has generally considered them to be stem or crown group members of Mandibulata, due to the presence of Mandible (arthropod mouthpart), mandibles in well-preserved species. Taxonomy Hymenocarines are characterized by the combination of the following characters: bivalved, convex carapace covering Cephalothorax, cephalothoracic (combined head and thorax) region; cephalothorax bearing multisegmented antennae (though as an exception antennae are absent in ''Odaraia'') and rounded Mandible (arthropod mouthpart), mandibles and likely Maxilla (arthropod mouthpart), maxillae, post maxillae limbs with spiny, subdivided basis and Arthropod leg, endopods (lower, leg-like parts) with well-d ...
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Waptia
''Waptia'' is an extinct genus of marine arthropod from the Middle Cambrian of North America. It grew to a length of , and had a large bivalved carapace and a segmented body terminating into a pair of tail flaps. It was an active swimmer and likely a predator of soft-bodied prey. It is also one of the oldest animals with direct evidence of Parental investment, brood care. ''Waptia fieldensis'' is the only species classified under the genus ''Waptia'', and is known from the Burgess Shale ''Lagerstätte'' of British Columbia, Canada. Specimens of ''Waptia'' are also known from the Spence Shale of Utah, United States. Based on the number of individuals, ''Waptia fieldensis'' is the third most abundant arthropod from the Burgess Shale Formation, with thousands of specimens collected. It was among the first fossils found by the American paleontologist Charles D. Walcott in 1909. He described it in 1912 and named it after two mountains near the discovery site – Wapta Mountain and Moun ...
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Canadaspis
''Canadaspis'' ("Shield of Canada") is an extinct genus of bivalved Cambrian marine arthropod, known from North America and China. They are thought to have been benthic feeders that moved mainly by walking and possibly used its biramous appendages to stir mud in search of food. They have been placed within the Hymenocarina, which includes other bivalved Cambrian arthropods. Description ''Canadaspis'' ''perfecta'' The bivalved carapaces of ''Canadaspis'' ''perfecta'' are typically in length, which taper towards the front end. The head had a small pair of eyes borne on short stalks. Between the eyes is a forward pointing spine, as well as a pair of short antennae, which appear to lack segmentation. Similar antennae are known from '' Waptia'', and are probably homologous to the hemi-ellipsoid bodies of crustaceans, and thus likely have an olfactory function. The head also has another pair of larger, segmented antennae, probably with more than 12 segments, the segments increased ...
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Odaraia
''Odaraia'' is an extinct genus of bivalved Hymenocarina, hymenocarine arthropod with a single known species ''Odaraia alata'', found in the Middle Cambrian Burgess Shale in British Columbia, Canada. History of research and taxonomy The genus and species were first described by Charles Doolittle Walcott, Charles Walcott in 1912. It was placed into its own family, Odaraiidae by Simonetta and Delle Cave in 1975,Simonetta AM, Delle Cave L . 1975 The Cambrian non-trilobite arthropods from the Burgess Shale of British Columbia: a study of their comparative morphology, taxonomy and evolutionary significance. ''Palaeontogr. Ital.'' which has been used to include other genera of bivalved Cambrian arthropods. The species was redescribed by Derek Briggs 1981, and again redescribed in 2024 by Izquierdo-López and Caron. While originally ''Odaraia'' and other odaraiids have been interpreted as Basal (phylogenetics), basal euarthropods not closely related to any living arthropod group, the co ...
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Arthropods
Arthropods ( ) are invertebrates in the phylum Arthropoda. They possess an arthropod exoskeleton, exoskeleton with a cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate, a body with differentiated (Metamerism (biology), metameric) Segmentation (biology), segments, and paired jointed appendages. In order to keep growing, they must go through stages of moulting, a process by which they shed their exoskeleton to reveal a new one. They form an extremely diverse group of up to ten million species. Haemolymph is the analogue of blood for most arthropods. An arthropod has an open circulatory system, with a body cavity called a haemocoel through which haemolymph circulates to the interior Organ (anatomy), organs. Like their exteriors, the internal organs of arthropods are generally built of repeated segments. They have ladder-like nervous systems, with paired Anatomical terms of location#Dorsal and ventral, ventral Ventral nerve cord, nerve cord ...
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Mandibulata
The clade Mandibulata constitutes one of the major subdivisions of the phylum Arthropoda, alongside Chelicerata. Mandibulates include the crustaceans, myriapods (centipedes and millipedes, among others), and all true insects. The name "Mandibulata" refers to the mandibles, a modified pair of limbs used in food processing, the presence of which are characteristic of most members of the group. The mandibulates are divided between the extant groups Myriapoda (millipedes and centipedes, among others) and Pancrustacea (including crustaceans and hexapods, the latter group containing insects). Molecular phylogenetic studies suggest that the living arthropods are related as shown in the cladogram below. Crustaceans do not form a monophyletic In biological cladistics for the classification of organisms, monophyly is the condition of a taxonomic grouping being a clade – that is, a grouping of organisms which meets these criteria: # the grouping contains its own most recent ...
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Isoxyida
Isoxyids are members of the order Isoxyida and the family Isoxyidae, a group of basal arthropods that existed during the Cambrian period. It contains two genera, ''Isoxys'', with 20 species found worldwide, and ''Surusicaris'' known from a single species found in the Burgess Shale of Canada. They are distinguished by their bivalved carapaces and pair of upward curving grasping frontal appendages. Description Isoxyids have a combination of features seen in both stem-group arthropods, as well as more advanced taxa. Isoxyids have semicircular bivalved Carapace, carapaces, large, spherical Arthropod eye, eyes, a pair of large upward curling frontal appenages covered in spines, and pairs of biramous limbs running along the body. The trunk region appears to be unsegmented and lacks sclerotisation, similar to Radiodonta, radiodonts, with a segmented and sclerotised (arthropodized) trunk being characteristic of most arthropods. Currently only two genera have been accepted to be members ...
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Cambrian
The Cambrian ( ) is the first geological period of the Paleozoic Era, and the Phanerozoic Eon. The Cambrian lasted 51.95 million years from the end of the preceding Ediacaran period 538.8 Ma (million years ago) to the beginning of the Ordovician Period 486.85 Ma. Most of the continents lay in the southern hemisphere surrounded by the vast Panthalassa Ocean. The assembly of Gondwana during the Ediacaran and early Cambrian led to the development of new convergent plate boundaries and continental-margin arc magmatism along its margins that helped drive up global temperatures. Laurentia lay across the equator, separated from Gondwana by the opening Iapetus Ocean. The Cambrian marked a profound change in life on Earth; prior to the Period, the majority of living organisms were small, unicellular and poorly preserved. Complex, multicellular organisms gradually became more common during the Ediacaran, but it was not until the Cambrian that fossil diversity seems to rapidly ...
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Chuandianella
''Chuandianella ovata'' is an extinct bivalved arthropod that lived during Cambrian Stage 3 of the Early Cambrian (about 520 to 516 million years ago). It is the only species classified under the genus ''Chuandianella''. Its fossils were recovered from the Chengjiang Biota in Yunnan, China. Taxonomy It was originally described in 1975 under the "ostracod"-like genus '' Mononotella'', as ''Mononotella ovata''. In 1991, the paleontologists Hou Xian-guang and Jan Bergström reclassified it under the new genus ''Chuandianella'' when additional discoveries of more complete specimens made its resemblance to '' Waptia fieldensis'' more apparent. In 2004, paleontologist Jun-Yuan Chen tentatively transferred it to the genus ''Waptia''. However, ''C. ovata'' had eight abdominal somites in contrast to five in ''W. fieldensis''. Its limbs were biramous and were undifferentiated, unlike those of ''W. fieldensis''. Other authors deemed these differences to be enough to separate it from ''W ...
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Pancrustacea
Pancrustacea is the clade that comprises all crustaceans and all hexapods (insects and relatives). This grouping is contrary to the Atelocerata hypothesis, in which Hexapoda and Myriapoda are sister taxa, and Crustacea are only more distantly related. As of 2010, the Pancrustacea taxon was considered well accepted, with most studies recovering Hexapoda within Crustacea. The clade has also been called Tetraconata, referring to having a four-part cone in the ommatidium. The term "Tetraconata" is preferred by some scientists in order to avoid confusion with the use of "pan-" to indicate a clade that includes a crown group and all of its stem group representatives. Molecular studies A monophyletic Pancrustacea has been supported by several molecular studies, in most of which the subphylum Crustacea is paraphyletic with regard to hexapods (that is, that hexapods, including insects, are derived from crustacean ancestors). This means that within Pancrustacea, only some members are ...
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Monophyletic
In biological cladistics for the classification of organisms, monophyly is the condition of a taxonomic grouping being a clade – that is, a grouping of organisms which meets these criteria: # the grouping contains its own most recent common ancestor (or more precisely an ancestral population), i.e. excludes non-descendants of that common ancestor # the grouping contains all the descendants of that common ancestor, without exception Monophyly is contrasted with paraphyly and polyphyly as shown in the second diagram. A ''paraphyletic'' grouping meets 1. but not 2., thus consisting of the descendants of a common ancestor, excepting one or more monophyletic subgroups. A '' polyphyletic'' grouping meets neither criterion, and instead serves to characterize convergent relationships of biological features rather than genetic relationships – for example, night-active primates, fruit trees, or aquatic insects. As such, these characteristic features of a polyphyletic grouping ...
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Paraphyletic
Paraphyly is a taxonomic term describing a grouping that consists of the grouping's last common ancestor and some but not all of its descendant lineages. The grouping is said to be paraphyletic ''with respect to'' the excluded subgroups. In contrast, a monophyletic grouping (a clade) includes a common ancestor and ''all'' of its descendants. The terms are commonly used in phylogenetics (a subfield of biology) and in the tree model of historical linguistics. Paraphyletic groups are identified by a combination of synapomorphies and symplesiomorphies. If many subgroups are missing from the named group, it is said to be polyparaphyletic. The term received currency during the debates of the 1960s and 1970s accompanying the rise of cladistics, having been coined by zoologist Willi Hennig to apply to well-known taxa like Reptilia (reptiles), which is paraphyletic with respect to birds. Reptilia contains the last common ancestor of reptiles and all descendants of that ancestor exc ...
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Polyphyletic
A polyphyletic group is an assemblage that includes organisms with mixed evolutionary origin but does not include their most recent common ancestor. The term is often applied to groups that share similar features known as Homoplasy, homoplasies, which are explained as a result of convergent evolution. The arrangement of the members of a polyphyletic group is called a polyphyly .. [Source for pronunciation.] It is contrasted with monophyly and paraphyly. For example, the biological characteristic of warm-bloodedness evolved separately in the ancestors of mammals and the ancestors of birds; "warm-blooded animals" is therefore a polyphyletic grouping. Other examples of polyphyletic groups are algae, C4 photosynthesis, C4 photosynthetic plants, and Xenarthra#Evolutionary relationships, edentates. Many taxonomists aim to avoid homoplasies in grouping taxa together, with a goal to identify and eliminate groups that are found to be polyphyletic. This is often the stimulus for major re ...
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