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Haptista
Haptista is a proposed group of protists made up of centrohelids and haptophytes. Phylogenomic studies indicate that Haptista, together with ''Ancoracysta twista'', forms a sister clade to the SAR supergroup#Internal phylogeny, TSAR Supergroup (biology), supergroup, but it may also be sister to the Cryptista (+Archaeplastida). It is thus one of the earliest diverging Diaphoretickes. Taxonomy Based on studies done by Cavalier-Smith, Chao & Lewis 2015 and Ruggiero et al. 2015. * Subphylum Centroheliozoa Cushman & Jarvis 1929 sensu Durrschmidt & Patterson 1987 [Heliozoa Haeckel 1862 stat. n. Margulis 1974 em. Cavalier-Smith 2003] ** Class Centrohelea Kuhn 1926 stat. n. Cavalier-Smith 1993 [Centroplastiales; Centrohelina Hartmann 1913; Centroplasthelida Febvre-Chevalier 1984] * Subphylum Haptophyte, Haptophytina Cavalier-Smith 2015 (Haptophyta Hibberd 1976 sensu Ruggerio et al. 2015) ** Clade Rappemonada Kim et al. 2011 *** Class Rappephyceae Cavalier-Smith 2015 ** Clade Haptomonada ( ...
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Protist
A protist ( ) or protoctist is any eukaryotic organism that is not an animal, land plant, or fungus. Protists do not form a natural group, or clade, but are a paraphyletic grouping of all descendants of the last eukaryotic common ancestor excluding land plants, animals, and fungi. Protists were historically regarded as a separate taxonomic kingdom known as Protista or Protoctista. With the advent of phylogenetic analysis and electron microscopy studies, the use of Protista as a formal taxon was gradually abandoned. In modern classifications, protists are spread across several eukaryotic clades called supergroups, such as Archaeplastida ( photoautotrophs that includes land plants), SAR, Obazoa (which includes fungi and animals), Amoebozoa and " Excavata". Protists represent an extremely large genetic and ecological diversity in all environments, including extreme habitats. Their diversity, larger than for all other eukaryotes, has only been discovered in rece ...
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Diaphoretickes
Diaphoretickes is a major group of eukaryote, eukaryotic organisms spanning over 400,000 species. The majority of the earth's biomass that carries out photosynthesis belongs to Diaphoretickes. In older classification systems, members of the Diaphoretickes were variously placed in the Kingdom (biology), kingdoms Protozoa or Protist, Protista. Etymology The name Diaphoretickes derives (''diaforetikés'') meaning diverse, dissimilar, referring to the wide morphology (biology), morphological and cellular diversity among members of this clade. History Eukaryotes, organisms whose cells contain a cell nucleus, nucleus, have been traditionally grouped into four kingdom (biology), kingdoms: animals, plants, fungi and protists. In the late 20th century, molecular phylogenetic analyses revealed that protists are a paraphyletic assortment of many independent evolutionary lineages or clades, from which animals, fungi and plants evolved. However, the relationships between these clades re ...
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Haptista
Haptista is a proposed group of protists made up of centrohelids and haptophytes. Phylogenomic studies indicate that Haptista, together with ''Ancoracysta twista'', forms a sister clade to the SAR supergroup#Internal phylogeny, TSAR Supergroup (biology), supergroup, but it may also be sister to the Cryptista (+Archaeplastida). It is thus one of the earliest diverging Diaphoretickes. Taxonomy Based on studies done by Cavalier-Smith, Chao & Lewis 2015 and Ruggiero et al. 2015. * Subphylum Centroheliozoa Cushman & Jarvis 1929 sensu Durrschmidt & Patterson 1987 [Heliozoa Haeckel 1862 stat. n. Margulis 1974 em. Cavalier-Smith 2003] ** Class Centrohelea Kuhn 1926 stat. n. Cavalier-Smith 1993 [Centroplastiales; Centrohelina Hartmann 1913; Centroplasthelida Febvre-Chevalier 1984] * Subphylum Haptophyte, Haptophytina Cavalier-Smith 2015 (Haptophyta Hibberd 1976 sensu Ruggerio et al. 2015) ** Clade Rappemonada Kim et al. 2011 *** Class Rappephyceae Cavalier-Smith 2015 ** Clade Haptomonada ( ...
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Centrohelid
The centrohelids or centroheliozoa are a large group of heliozoan protists. They include both mobile and sessile forms, found in freshwater and marine environments, especially at some depth. Characteristics Individuals are unicellular and spherical, usually around 30–80 μm in diameter, and covered with long radial axopods, narrow cellular projections that capture food and allow mobile forms to move about. A few genera have no cell covering, but most have a gelatinous coat holding scales and spines, produced in special deposition vesicles. These may be organic or siliceous and come in various shapes and sizes. For instance, in '' Raphidiophrys'' the coat extends along the bases of the axopods, covering them with curved spicules that give them a pine-treeish look, and in ''Raphidiocystis'' there are both short cup-shaped spicules and long tubular spicules that are only a little shorter than the axopods. Some other common genera include '' Heterophrys'', ''Actinocystis'', and ...
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Supergroup (biology)
A supergroup or super-group, in systematics, is a large group of organisms that monophyletic, share one common ancestor and have important defining characteristics. It is an informal, mostly arbitrary rank in biology, biological Taxonomy (biology), taxonomy that is often greater than phylum or kingdom (biology), kingdom, although some supergroups are also treated as phylum, phyla. Eukaryotic supergroups Since the decade of the 2000s, the Eukaryote#Phylogeny, eukaryotic tree of life (abbreviated as eToL) has been divided into 5–8 major groupings called 'supergroups'. These groupings were established after the idea that only monophyletic groups should be accepted as ranks, as an alternative to the use of paraphyletic kingdom Protista. In the early days of the eToL six traditional supergroups were considered: Amoebozoa, Opisthokonta, "Excavata", Archaeplastida, "Chromalveolata" and Rhizaria. Since then, the eToL has been rearranged profoundly, and most of these groups were found as pa ...
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Cryptista
Cryptista is a clade of alga-like eukaryotes. It is most likely related to Archaeplastida which includes plants and many algae, within the larger group Diaphoretickes. Other characteristic features of cryptophyte mtDNAs include large syntenic clusters resembling α-proteobacterial operons that encode bacteria-like rRNAs, tRNAs, and ribosomal protein genes. Additionally, they are an evolutionarily significant lineage found in mostly marine, glacial and freshwater environments. Although it has sometimes placed along with Haptista in the group Hacrobia, within the kingdom Chromista, most recent studies have found that Hacrobia is not a clade. For example, in 2016, a broad phylogenomic study found that cryptists fall within the group Archaeplastida, while haptophytes are closely related to the SAR supergroup. Taxonomy Based on studies done by Cavalier-Smith, Chao & Lewis 2015 * Corbihelia ** Clade Endohelia Cavalier-Smith 2015 *** Clade Endohelea Cavalier-Smith 2012 * Cla ...
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Archaeplastida
The Archaeplastida (or kingdom Plantae '' sensu lato'' "in a broad sense"; pronounced ) are a major group of eukaryotes, comprising the photoautotrophic red algae (Rhodophyta), green algae, land plants, and the minor group glaucophytes. It also includes the non-photosynthetic lineage Rhodelphidia, a predatorial (eukaryotrophic) flagellate that is sister to the Rhodophyta, and probably the microscopic picozoans. The Archaeplastida have chloroplasts that are surrounded by two membranes, suggesting that they were acquired directly through a single endosymbiosis event by phagocytosis of a cyanobacterium. All other groups which have chloroplasts, besides the amoeboid genus '' Paulinella'', have chloroplasts surrounded by three or four membranes, suggesting they were acquired secondarily from red or green algae. Unlike red and green algae, glaucophytes have never been involved in secondary endosymbiosis events. The cells of the Archaeplastida typically lack centrioles and have mit ...
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Centrohelida
The centrohelids or centroheliozoa are a large group of heliozoan protists. They include both mobile and sessile forms, found in freshwater and marine environments, especially at some depth. Characteristics Individuals are unicellular and spherical, usually around 30–80 μm in diameter, and covered with long radial axopods, narrow cellular projections that capture food and allow mobile forms to move about. A few genera have no cell covering, but most have a gelatinous coat holding scales and spines, produced in special deposition vesicles. These may be organic or siliceous and come in various shapes and sizes. For instance, in '' Raphidiophrys'' the coat extends along the bases of the axopods, covering them with curved spicules that give them a pine-treeish look, and in ''Raphidiocystis'' there are both short cup-shaped spicules and long tubular spicules that are only a little shorter than the axopods. Some other common genera include '' Heterophrys'', ''Actinocystis'', and ...
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Rappephyceae
Rappephyceae, or Rappemonads, are a small family of protists first described in 2011, of uncertain phylogenic affinity. It has been discussed as a possible member of a larger clade Haptophyta. This newly identified taxonomic class of phytoplankton are named after a professor from the Hawai’i institute of marine biology, known as Michael Rappé. Rappé discovered these phytoplankton in the Atlantic Ocean and published his findings on their DNA in 1998. Current research has shown that these organisms provide an immense amount of nutritional molecules, such as oxygen, for other organisms using biochemical processes like photosynthesis and carbon fixation. Classification Rappephyceae belong to the Haptophyte clade. The haptophytes comprise around 500 marine algal species. One of the most novel characteristics of haptophytes is their calcite, or calcium carbonate, scales that cover the cell. These are also known as coccoliths and the whole organism can be referred to as a coccol ...
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Prymnesiophyceae
Prymnesiophyceae is a haptophyte class. Although it was originally described by Casper in 1972, it did not receive a Latin diagnosis (a requirement for valid publication under the International Code of Botanical Nomenclature The ''International Code of Nomenclature for algae, fungi, and plants'' (ICN or ICNafp) is the set of rules and recommendations dealing with the formal botanical names that are given to plants, fungi and a few other groups of organisms, all tho ...) until Hibberd provided one in 1976. References Haptophyte classes Haptista classes {{Haptophyte-stub ...
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