Equidae
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Equidae
Equidae (commonly known as the horse family) is the Taxonomy (biology), taxonomic Family (biology), family of Wild horse, horses and related animals, including Asinus, asses, zebra, zebras, and many extinct species known only from fossils. The family evolved more than 50 million years ago, in the Eocene epoch, from a small, multi-toed ungulate into larger, single-toed animals. All Extant taxon, extant species are in the genus ''Equus (genus), Equus'', which originated in North America. Equidae belongs to the order Perissodactyla, which includes the extant tapirs and rhinoceros, and several extinct families. It is more specifically grouped within the superfamily (taxonomy), superfamily Equoidea, the only other family being the extinct Palaeotheriidae. The term equid refers to any member of this family, including any equinae, equine. Evolution The oldest known fossils assigned to Equidae were found in North America, and date from the early Eocene epoch, 54 million years ago. Th ...
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Early Eocene
In the geologic timescale the Ypresian is the oldest age (geology), age or lowest stage (stratigraphy), stratigraphic stage of the Eocene. It spans the time between , is preceded by the Thanetian Age (part of the Paleocene) and is followed by the Eocene Lutetian Age. The Ypresian is consistent with the Lower Eocene (Early Eocene). Events The Ypresian Age begins during the throes of the Paleocene–Eocene Thermal Maximum (PETM). The Fur Formation in Denmark, the Messel shales in Germany, the Oise amber of France and Cambay amber of India are of this age. The Eocene Okanagan Highlands are an uplands subtropical to temperate series of lakes from the Ypresian. The Ypresian is additionally marked by another warming event called the Early Eocene Climatic Optimum (EECO). The EECO is the longest sustained warming event in the Cenozoic record, lasting about 2–3 million years between 53 and 50 Ma. The interval is characterized by low oxygen-18 isotopes, high levels of atmospheric pCO2 ...
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Genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In binomial nomenclature, the genus name forms the first part of the binomial species name for each species within the genus. :E.g. ''Panthera leo'' (lion) and ''Panthera onca'' (jaguar) are two species within the genus ''Panthera''. ''Panthera'' is a genus within the family Felidae. The composition of a genus is determined by taxonomy (biology), taxonomists. The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including the idea that a newly defined genus should fulfill these three criteria to be descriptively useful: # monophyly – all descendants of an ancestral taxon are grouped together (i.e. Phylogeneti ...
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Hindgut Fermentation
Hindgut fermentation is a digestive process seen in monogastric herbivores (animals with a simple, single-chambered stomach). Cellulose is digested with the aid of symbiotic microbes including bacteria, archaea, and eukaryotes. The microbial fermentation occurs in the digestive organs that follow the small intestine: the cecum and large intestine. Examples of hindgut fermenters include proboscideans and large odd-toed ungulates such as horses and rhinos, as well as small animals such as rodents, rabbits and koalas. In contrast, foregut fermentation is the form of cellulose digestion seen in ruminants such as cattle which have a four-chambered stomach, as well as in sloths, macropodids, some monkeys, and one bird, the hoatzin.. Cecum Hindgut fermenters generally have a cecum and large intestine that are much larger and more complex than those of a foregut or midgut fermenter.
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Palaeotheriidae
Palaeotheriidae is an extinct family of herbivorous perissodactyl mammals that inhabited Europe, with less abundant remains also known from Asia, from the mid-Eocene to the early Oligocene. They are classified in Equoidea, along with the living family Equidae (which includes zebras, horses and asses). Morphology Palaeotheres ranged widely in size, from small species like '' Palaeotherium lautricense,'' which is estimated to have only weighed to large species like '' Palaeotherium magnum'', which are comparable in size to living equines, with body masses over . Their teeth are brachydont (low crowned). According to Danilo et al. 2013., paleotheriids are distinguished from other equoids by one unambiguous synapomorphy "the nasal notch opening distally to the canine, above the postcanine diastema" and two unambiguous character state changes "an average metaconule on he fourth premolar and "an oblique metastyle on he first and second molars. Taxonomy Palaeotheriidae is ge ...
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Type Species
In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological Type (biology), type wiktionary:en:specimen, specimen (or specimens). Article 67.1 A similar concept is used for suprageneric groups and called a type genus. In botanical nomenclature, these terms have no formal standing under the International Code of Nomenclature for algae, fungi, and plants, code of nomenclature, but are sometimes borrowed from zoological nomenclature. In botany, the type of a genus name is a specimen (or, rarely, an illustration) which is also the type of a species name. The species name with that type can also be referred to as the type of the genus name. Names of genus and family ranks, the various subdivisions of those ranks, and some higher-rank names based on genus names, have suc ...
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Hyracotherium
''Hyracotherium'' ( ; "hyrax-like beast") is an extinction, extinct genus of small (about 60 cm in length) perissodactyl ungulates that was found in the London Clay formation. This small, fox-sized animal is (for some scientists) considered to be the earliest known member of Equidae before the type species, ''H. leporinum'', was reclassified as a palaeothere, a perissodactyl family (biology), family related to both horses and brontotheres. The remaining species are now thought to belong to different genera, such as ''Eohippus'', which had previously been synonymised with ''Hyracotherium''. Description ''Hyracotherium'' averaged 78 cm (2.5 feet) in length and weighed about 9 kg (20 pounds). It had a short face with eye sockets in the middle and a short diastema (the space between the front teeth and the cheek teeth). The skull was long, having 44 low-crowned teeth. Although it had low-crowned teeth, the beginnings of the characteristic horse-like ridges on the molars ...
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Extinct Horses
Extinction is the termination of an organism by the death of its last member. A taxon may become functionally extinct before the death of its last member if it loses the capacity to reproduce and recover. As a species' potential range may be very large, determining this moment is difficult, and is usually done retrospectively. This difficulty leads to phenomena such as Lazarus taxa, where a species presumed extinct abruptly "reappears" (typically in the fossil record) after a period of apparent absence. Over five billion species are estimated to have died out. It is estimated that there are currently around 8.7 million species of eukaryotes globally, possibly many times more if microorganisms are included. Notable extinct animal species include non-avian dinosaurs, saber-toothed cats, and mammoths. Through evolution, species arise through the process of speciation. Species become extinct when they are no longer able to survive in changing conditions or against superi ...
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Equinae
Equinae is a subfamily of the family Equidae, known from the Hemingfordian stage of the Early Miocene (16 million years ago) onwards. They originated in North America, before dispersing to every continent except Australia and Antarctica. They are thought to be a monophyletic grouping. Members of the subfamily are referred to as equines; the only extant equines are the horses, asses, and zebra Zebras (, ) (subgenus ''Hippotigris'') are African equines with distinctive black-and-white striped coats. There are three living species: Grévy's zebra (''Equus grevyi''), the plains zebra (''E. quagga''), and the mountain zebra (''E. ...s of the genus ''Equus'', with two other genera '' Haringtonhippus'' and '' Hippidion'' becoming extinct at the beginning of the Holocene, around 11–12,000 years ago. The subfamily contains two tribes, the Equini and the Hipparionini, as well as two unplaced genera, '' Merychippus'' and '' Scaphohippus''. Members of the family ancestrally ...
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Palaeotheriidae
Palaeotheriidae is an extinct family of herbivorous perissodactyl mammals that inhabited Europe, with less abundant remains also known from Asia, from the mid-Eocene to the early Oligocene. They are classified in Equoidea, along with the living family Equidae (which includes zebras, horses and asses). Morphology Palaeotheres ranged widely in size, from small species like '' Palaeotherium lautricense,'' which is estimated to have only weighed to large species like '' Palaeotherium magnum'', which are comparable in size to living equines, with body masses over . Their teeth are brachydont (low crowned). According to Danilo et al. 2013., paleotheriids are distinguished from other equoids by one unambiguous synapomorphy "the nasal notch opening distally to the canine, above the postcanine diastema" and two unambiguous character state changes "an average metaconule on he fourth premolar and "an oblique metastyle on he first and second molars. Taxonomy Palaeotheriidae is ge ...
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Equoidea
Equoidea is a superfamily of hippomorph perissodactyls containing the Equidae, Palaeotheriidae, and other basal equoids of unclear affinities, of which members of the genus '' Equus'' are the only extant species. The earliest fossil record of the Equoidea proves unclear, but they possibly could have originated during the late Paleocene in Europe or Asia. Definite fossil records of equoids are recorded by the earliest Eocene, in which the earliest equids in North America and basal equoids of unclear affinities in Europe both appeared. Palaeotheres are thought to have originated later in the early Eocene of Europe, although some researchers disagree on whether the subfamily Pachynolophinae belongs to the Palaeotheriidae. Equoids may in part be defined by dental synapomorphies from its molars The molars or molar teeth are large, flat tooth, teeth at the back of the mouth. They are more developed in mammal, mammals. They are used primarily to comminution, grind food during mast ...
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Superfamily (taxonomy)
In biology, taxonomic rank (which some authors prefer to call nomenclatural rank because ranking is part of nomenclature rather than taxonomy proper, according to some definitions of these terms) is the relative or absolute level of a group of organisms (a ''taxon'') in a hierarchy that reflects evolutionary relationships. Thus, the most inclusive clades (such as Eukarya and Animalia) have the highest ranks, whereas the least inclusive ones (such as ''Homo sapiens'' or ''Bufo bufo'') have the lowest ranks. Ranks can be either relative and be denoted by an indented taxonomy in which the level of indentation reflects the rank, or absolute, in which various terms, such as species, genus, family, order, class, phylum, kingdom, and domain designate rank. This page emphasizes absolute ranks and the rank-based codes (the Zoological Code, the Botanical Code, the Code for Cultivated Plants, the Prokaryotic Code, and thCode for Viruses require them. However, absolute ranks are not req ...
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Rhinoceros
A rhinoceros ( ; ; ; : rhinoceros or rhinoceroses), commonly abbreviated to rhino, is a member of any of the five extant taxon, extant species (or numerous extinct species) of odd-toed ungulates (perissodactyls) in the family (biology), family Rhinocerotidae; it can also refer to a member of any of the extinct species of the superfamily Rhinocerotoidea. Two of the extant species are native to Africa, and three to South Asia, South and Southeast Asia. Rhinoceroses are some of the largest remaining megafauna: all weigh over half a tonne in adulthood. They have a herbivore, herbivorous diet, small brains for mammals of their size, one or two horns, and a thick , protective skin formed from layers of collagen positioned in a crystal structure, lattice structure. They generally eat leafy material, although their ability to ferment food in their colon (anatomy), hindgut allows them to subsist on more fibrous plant matter when necessary. Unlike other perissodactyls, the two African ...
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